Department of Ecology and Evolutionary Biology, University of Kansas, Lawrence, KS, USA, Heidelberg Institute for Theoretical Studies, Heidelberg 69118, Germany.|Department of Ecology and Evolutionary Biology, University of Michigan, Ann Arbor, MI, USA.|Interrobang Corporation, Wake Forest, NC 27587, USA and.|Department of Ecology and Evolutionary Biology, University of Michigan, Ann Arbor, MI, USA.|National Evolutionary Synthesis Center, Duke University, Durham, NC, USA.|Department of Ecology and Evolutionary Biology, University of Kansas, Lawrence, KS, USA, Heidelberg Institute for Theoretical Studies, Heidelberg 69118, Germany.|National Evolutionary Synthesis Center, Duke University, Durham, NC, USA.|Department of Ecology and Evolutionary Biology, University of Michigan, Ann Arbor, MI, USA.