Department of Ecology and Evolutionary Biology, University of California, Irvine, California, United States of America, Department of Mathematics, Rowland Hall, University of California, Irvine, California, United States of America.|Department of Mathematics, Rowland Hall, University of California, Irvine, California, United States of America.|Department of Ecology and Evolutionary Biology, University of California, Irvine, California, United States of America, Department of Mathematics, Rowland Hall, University of California, Irvine, California, United States of America.