Computational Biology Program, Oregon Health and Science University, Portland, OR 97239, USA.|Division of Human Biology, Fred Hutchinson Cancer Research Center, Seattle, WA 98109, USA.|Division of Human Biology, Fred Hutchinson Cancer Research Center, Seattle, WA 98109, USA.|Computational Biology Program, Oregon Health and Science University, Portland, OR 97239, USA.|Computational Biology Program, Oregon Health and Science University, Portland, OR 97239, USA.