School of Mathematics and Physics, The University of Queensland, Brisbane 4066, Australia, ARC Centre of Excellence for Mathematical and Statistical Frontiers, QUT, Brisbane, Australia.|CREST, ENSAI, Campus de Ker-Lann, Rue Blaise Pascal, BP 37203, 35172 Bruz cedex, France.|Inria, SISTM, Talence and Inserm, U897, Bordeaux, Bordeaux University, Bordeaux and Vaccine Research Institute, Creteil, France.|Inria, SISTM, Talence and Inserm, U897, Bordeaux, Bordeaux University, Bordeaux and Vaccine Research Institute, Creteil, France.