School of Botany and Zoology, Faculty of Science, Australian National University, ACT 0200, Australia. mark.gibbs@anu.edu.au|School of Botany and Zoology, Faculty of Science, Australian National University, ACT 0200, Australia. mark.gibbs@anu.edu.au|School of Botany and Zoology, Faculty of Science, Australian National University, ACT 0200, Australia. mark.gibbs@anu.edu.au