Department of Biology, University of Nebraska, Omaha, NE 68182, USA. glu3@mail.unomaha.edu|Department of Biology, University of Nebraska, Omaha, NE 68182, USA. glu3@mail.unomaha.edu|Department of Biology, University of Nebraska, Omaha, NE 68182, USA. glu3@mail.unomaha.edu|Department of Biology, University of Nebraska, Omaha, NE 68182, USA. glu3@mail.unomaha.edu