Department of Developmental, Molecular & Chemical Biology. Sackler School of Graduate Biomedical Sciences, Tufts University School of Medicine, Boston, Massachusetts, United States of America.|Department of Developmental, Molecular & Chemical Biology. Sackler School of Graduate Biomedical Sciences, Tufts University School of Medicine, Boston, Massachusetts, United States of America.|Department of Developmental, Molecular & Chemical Biology. Sackler School of Graduate Biomedical Sciences, Tufts University School of Medicine, Boston, Massachusetts, United States of America.|Department of Developmental, Molecular & Chemical Biology. Sackler School of Graduate Biomedical Sciences, Tufts University School of Medicine, Boston, Massachusetts, United States of America.|Department of Biology, Boston College, Boston, Massachusetts, United States of America.|Department of Developmental, Molecular & Chemical Biology. Sackler School of Graduate Biomedical Sciences, Tufts University School of Medicine, Boston, Massachusetts, United States of America.