Kavli Institute for Systems Neuroscience and Centre for Neural Computation, NTNU, Trondheim, Norway.|Kavli Institute for Systems Neuroscience and Centre for Neural Computation, NTNU, Trondheim, Norway.|Kavli Institute for Systems Neuroscience and Centre for Neural Computation, NTNU, Trondheim, Norway, Nordita, KTH and Stockholm University, Stockholm, Sweden.