Zeeman Institute for Systems Biology & Infectious Disease Epidemiology Research, University of Warwick, Coventry, United Kingdom, Mathematics Institute, University of Warwick, Coventry, United Kingdom.|Department of Statistics, University of Warwick, Coventry, United Kingdom.|Faculty of Biology, Medicine & Health, University of Manchester, Manchester, United Kingdom.|Systems Microscopy Centre, University of Manchester, Manchester, United Kingdom.|Faculty of Biology, Medicine & Health, University of Manchester, Manchester, United Kingdom.|Systems Microscopy Centre, University of Manchester, Manchester, United Kingdom.|Department of Physiology, Anatomy and Genetics, University of Oxford, Oxford, United Kingdom.|Systems Microscopy Centre, University of Manchester, Manchester, United Kingdom.|Faculty of Biology, Medicine & Health, University of Manchester, Manchester, United Kingdom.|Department of Statistics, University of Warwick, Coventry, United Kingdom.|Zeeman Institute for Systems Biology & Infectious Disease Epidemiology Research, University of Warwick, Coventry, United Kingdom, Mathematics Institute, University of Warwick, Coventry, United Kingdom.