Department of Neurobiology, Harvard Medical School, 220 Longwood Avenue, Boston, MA 02115, USA.|Department of Neurobiology, Harvard Medical School, 220 Longwood Avenue, Boston, MA 02115, USA.|Department of Neurobiology, Harvard Medical School, 220 Longwood Avenue, Boston, MA 02115, USA.|Department of Neurobiology, Harvard Medical School, 220 Longwood Avenue, Boston, MA 02115, USA.