Genetics Program, Michigan State University, East Lansing, Michigan, United States of America.|Department of Plant Biology, Michigan State University, East Lansing, Michigan, United States of America.|Department of Plant Biology, Michigan State University, East Lansing, Michigan, United States of America.|Department of Plant Biology, Michigan State University, East Lansing, Michigan, United States of America.|Genetics Program, Michigan State University, East Lansing, Michigan, United States of America.