The Research School of Biology, Australian National University, Canberra ACT 2601, Australia.|The Research School of Biology, Australian National University, Canberra ACT 2601, Australia.|The Research School of Biology, Australian National University, Canberra ACT 2601, Australia.|The Research School of Biology, Australian National University, Canberra ACT 2601, Australia.|The Research School of Biology, Australian National University, Canberra ACT 2601, Australia.