Department of Life Sciences, University of Warwick, Coventry, United Kingdom.|CHICAS, Lancaster University, Bailrigg, Lancaster, United Kingdom.|Department of Infectious Disease Epidemiology, School of Public Health, Faculty of Medicine, St Marys Campus, Imperial College London, London, United Kingdom.|Department of Biostatistics, Vanderbilt University, Nashville, Tennessee, United States of America.|Center for Animal Disease Control, University of Miyazaki, Miyazaki, Japan.|US Geological Survey, Patuxent Wildlife Research Center, Laurel, Maryland, United States of America.|Center for Animal Disease Control, University of Miyazaki, Miyazaki, Japan.|Center for Infectious Disease Dynamics, Department of Biology, Eberly College of Science, The Pennsylvania State University, Pennsylvania, United States of America.|Department of Life Sciences, University of Warwick, Coventry, United Kingdom.|Center for Infectious Disease Dynamics, Department of Biology, Eberly College of Science, The Pennsylvania State University, Pennsylvania, United States of America.|Department of Life Sciences, University of Warwick, Coventry, United Kingdom.