Institute of Healthy Ageing, Department of Genetics, Evolution and Environment, University College London, London, United Kingdom.|European Molecular Biology Laboratory, European Bioinformatics Institute, Wellcome Genome Campus, Hinxton, Cambridge, United Kingdom.|Institute of Healthy Ageing, Department of Genetics, Evolution and Environment, University College London, London, United Kingdom.|Institute of Healthy Ageing, Department of Genetics, Evolution and Environment, University College London, London, United Kingdom.|European Molecular Biology Laboratory, European Bioinformatics Institute, Wellcome Genome Campus, Hinxton, Cambridge, United Kingdom.|Institute of Healthy Ageing, Department of Genetics, Evolution and Environment, University College London, London, United Kingdom.