Department of Biology, University of Virginia, Charlottesville, VA 22904, USA. pr8y@virginia.edu|Department of Biology, University of Virginia, Charlottesville, VA 22904, USA. pr8y@virginia.edu|Department of Biology, University of Virginia, Charlottesville, VA 22904, USA. pr8y@virginia.edu|Department of Biology, University of Virginia, Charlottesville, VA 22904, USA. pr8y@virginia.edu|Department of Biology, University of Virginia, Charlottesville, VA 22904, USA. pr8y@virginia.edu|Department of Biology, University of Virginia, Charlottesville, VA 22904, USA. pr8y@virginia.edu