School of Biological and Chemical Sciences, University of Exeter, Washington Singer Laboratories, Exeter, Devon, EX4 4PS, UK.|School of Biological and Chemical Sciences, University of Exeter, Washington Singer Laboratories, Exeter, Devon, EX4 4PS, UK.|School of Biological and Chemical Sciences, University of Exeter, Washington Singer Laboratories, Exeter, Devon, EX4 4PS, UK.|School of Biological and Chemical Sciences, University of Exeter, Washington Singer Laboratories, Exeter, Devon, EX4 4PS, UK.