The Institute for Molecular Bioscience, University of Queensland, Brisbane 4072, Australia. a.forrest@imb.uq.edu.au|The Institute for Molecular Bioscience, University of Queensland, Brisbane 4072, Australia. a.forrest@imb.uq.edu.au|The Institute for Molecular Bioscience, University of Queensland, Brisbane 4072, Australia. a.forrest@imb.uq.edu.au|The Institute for Molecular Bioscience, University of Queensland, Brisbane 4072, Australia. a.forrest@imb.uq.edu.au|The Institute for Molecular Bioscience, University of Queensland, Brisbane 4072, Australia. a.forrest@imb.uq.edu.au|The Institute for Molecular Bioscience, University of Queensland, Brisbane 4072, Australia. a.forrest@imb.uq.edu.au|The Institute for Molecular Bioscience, University of Queensland, Brisbane 4072, Australia. a.forrest@imb.uq.edu.au|The Institute for Molecular Bioscience, University of Queensland, Brisbane 4072, Australia. a.forrest@imb.uq.edu.au|The Institute for Molecular Bioscience, University of Queensland, Brisbane 4072, Australia. a.forrest@imb.uq.edu.au|The Institute for Molecular Bioscience, University of Queensland, Brisbane 4072, Australia. a.forrest@imb.uq.edu.au|The Institute for Molecular Bioscience, University of Queensland, Brisbane 4072, Australia. a.forrest@imb.uq.edu.au