Institute for Molecular Bioscience, Division of Cell Biology and Molecular Medicine, The University of Queensland, St. Lucia, Brisbane, Queensland, Australia.|Institute for Molecular Bioscience, Division of Cell Biology and Molecular Medicine, The University of Queensland, St. Lucia, Brisbane, Queensland, Australia.|Institute for Molecular Bioscience, Division of Cell Biology and Molecular Medicine, The University of Queensland, St. Lucia, Brisbane, Queensland, Australia.|Institute for Molecular Bioscience, Division of Cell Biology and Molecular Medicine, The University of Queensland, St. Lucia, Brisbane, Queensland, Australia.|Department of Anatomy and Cell Biology, University of Kansas Medical Center, Kansas City, Kansas, United States of America.|Institute for Molecular Bioscience, Division of Cell Biology and Molecular Medicine, The University of Queensland, St. Lucia, Brisbane, Queensland, Australia.|School of Mathematics and Physics, The University of Queensland, St. Lucia, Brisbane, Queensland, Australia.