Institute of Integrative Biology, University of Liverpool, Liverpool, UK. drigden@liv.ac.uk.|Institute of Integrative Biology, University of Liverpool, Liverpool, UK. drigden@liv.ac.uk.|Institute of Integrative Biology, University of Liverpool, Liverpool, UK. drigden@liv.ac.uk.|Institute of Integrative Biology, University of Liverpool, Liverpool, UK. drigden@liv.ac.uk.|Institute of Integrative Biology, University of Liverpool, Liverpool, UK. drigden@liv.ac.uk.|Institute of Integrative Biology, University of Liverpool, Liverpool, UK. drigden@liv.ac.uk.