Department of Ecology and Evolutionary Biology, Princeton University, Princeton, New Jersey, United States of America.|Department of Biological Sciences, Dartmouth College, Hanover, New Hampshire, United States of America.|Max Planck Institute for Terrestrial Microbiology, Marburg, Germany.|Max Planck Institute for Terrestrial Microbiology, Marburg, Germany.|Marine Population Modeling Group, Department of Mathematics and Statistics, University of Strathclyde, Glasgow, Scotland, United Kingdom.