Department of Biology, Boston College, Chestnut Hill, MA 02467, USA. barnetde@bc.edu|Department of Biology, Boston College, Chestnut Hill, MA 02467, USA. barnetde@bc.edu|Department of Biology, Boston College, Chestnut Hill, MA 02467, USA. barnetde@bc.edu|Department of Biology, Boston College, Chestnut Hill, MA 02467, USA. barnetde@bc.edu|Department of Biology, Boston College, Chestnut Hill, MA 02467, USA. barnetde@bc.edu