RIKEN Plant Science Center, Yokohama, Kanagawa, Japan. ryoko@psc.riken.jp|RIKEN Plant Science Center, Yokohama, Kanagawa, Japan. ryoko@psc.riken.jp|RIKEN Plant Science Center, Yokohama, Kanagawa, Japan. ryoko@psc.riken.jp|RIKEN Plant Science Center, Yokohama, Kanagawa, Japan. ryoko@psc.riken.jp|RIKEN Plant Science Center, Yokohama, Kanagawa, Japan. ryoko@psc.riken.jp|RIKEN Plant Science Center, Yokohama, Kanagawa, Japan. ryoko@psc.riken.jp