Joint Carnegie Mellon University-University of Pittsburgh Ph.D. Program in Computational Biology, Department of Computational and Systems Biology, University of Pittsburgh, PA 15260, USA.|Joint Carnegie Mellon University-University of Pittsburgh Ph.D. Program in Computational Biology, Department of Computational and Systems Biology, University of Pittsburgh, PA 15260, USA.|Joint Carnegie Mellon University-University of Pittsburgh Ph.D. Program in Computational Biology, Department of Computational and Systems Biology, University of Pittsburgh, PA 15260, USA.|Joint Carnegie Mellon University-University of Pittsburgh Ph.D. Program in Computational Biology, Department of Computational and Systems Biology, University of Pittsburgh, PA 15260, USA.|Joint Carnegie Mellon University-University of Pittsburgh Ph.D. Program in Computational Biology, Department of Computational and Systems Biology, University of Pittsburgh, PA 15260, USA.