Neuroscience Graduate Program, University of Washington, Seattle, Washington, United States of America.|Division of Biology and Bioengineering, California Institute of Technology, Pasadena, California, United States of America.|Division of Biology and Bioengineering, California Institute of Technology, Pasadena, California, United States of America.|Department of Biology, University of Washington, Seattle, Washington, United States of America.|Department of Physiology and Biophysics, University of Washington, Seattle, Washington, United States of America.