Institute of Molecular and Cellular Biology, Department of Life Science, National Tsing Hua University, Hsinchu, 30013, ROC, Taiwan. d898209@life.nthu.edu.tw|Institute of Molecular and Cellular Biology, Department of Life Science, National Tsing Hua University, Hsinchu, 30013, ROC, Taiwan. d898209@life.nthu.edu.tw|Institute of Molecular and Cellular Biology, Department of Life Science, National Tsing Hua University, Hsinchu, 30013, ROC, Taiwan. d898209@life.nthu.edu.tw|Institute of Molecular and Cellular Biology, Department of Life Science, National Tsing Hua University, Hsinchu, 30013, ROC, Taiwan. d898209@life.nthu.edu.tw|Institute of Molecular and Cellular Biology, Department of Life Science, National Tsing Hua University, Hsinchu, 30013, ROC, Taiwan. d898209@life.nthu.edu.tw|Institute of Molecular and Cellular Biology, Department of Life Science, National Tsing Hua University, Hsinchu, 30013, ROC, Taiwan. d898209@life.nthu.edu.tw|Institute of Molecular and Cellular Biology, Department of Life Science, National Tsing Hua University, Hsinchu, 30013, ROC, Taiwan. d898209@life.nthu.edu.tw|Institute of Molecular and Cellular Biology, Department of Life Science, National Tsing Hua University, Hsinchu, 30013, ROC, Taiwan. d898209@life.nthu.edu.tw|Institute of Molecular and Cellular Biology, Department of Life Science, National Tsing Hua University, Hsinchu, 30013, ROC, Taiwan. d898209@life.nthu.edu.tw