Evolutionary Biology Group, School of Biological, Biomedical & Environmental Sciences, The University of Hull, Hull, United Kingdom.|Evolutionary Biology Group, School of Biological, Biomedical & Environmental Sciences, The University of Hull, Hull, United Kingdom.|Institute of Evolutionary Biology, The University of Edinburgh, Edinburgh, United Kingdom.|Evolutionary Biology Group, School of Biological, Biomedical & Environmental Sciences, The University of Hull, Hull, United Kingdom.