Department of Plant Sciences, University of Tennessee, Knoxville, TN 37996, USA. syuan@utk.edu|Department of Plant Sciences, University of Tennessee, Knoxville, TN 37996, USA. syuan@utk.edu|Department of Plant Sciences, University of Tennessee, Knoxville, TN 37996, USA. syuan@utk.edu|Department of Plant Sciences, University of Tennessee, Knoxville, TN 37996, USA. syuan@utk.edu