The Peter Medawar Building for Pathogen Research and Department of Zoology, University of Oxford, OX1 3SY, UK. keith.jolley@medawar.ox.ac.uk|The Peter Medawar Building for Pathogen Research and Department of Zoology, University of Oxford, OX1 3SY, UK. keith.jolley@medawar.ox.ac.uk|The Peter Medawar Building for Pathogen Research and Department of Zoology, University of Oxford, OX1 3SY, UK. keith.jolley@medawar.ox.ac.uk