Department of Physics, Chemistry and Biology, Linköping University, Linköping, Sweden, Department of Biology, Colorado State University, Fort Collins, Colorado, United States of America, US National Institute of Health, Bethesda, Maryland, United States of America, University of Exeter, Exeter, United Kingdom.|US National Institute of Health, Bethesda, Maryland, United States of America, School of Veterinary Medicine and Science, University of Nottingham, Leicestershire, United Kingdom.|Department of Biology, Colorado State University, Fort Collins, Colorado, United States of America, US National Institute of Health, Bethesda, Maryland, United States of America.