Department of Ecology and Evolutionary Biology and Program in Computational Biology and Bioinformatics, Yale University, New Haven, CT 06520, USA.|Department of Ecology and Evolutionary Biology and Program in Computational Biology and Bioinformatics, Yale University, New Haven, CT 06520, USA.|Department of Ecology and Evolutionary Biology and Program in Computational Biology and Bioinformatics, Yale University, New Haven, CT 06520, USA.