Committee on Evolutionary Biology, University of Chicago, 1025 E. 57th St. Chicago, IL 60637, USA and Botany Department, Field Museum of Natural History, 1400 S. Lake Shore Dr. Chicago, IL 60605, USACommittee on Evolutionary Biology, University of Chicago, 1025 E. 57th St. Chicago, IL 60637, USA and Botany Department, Field Museum of Natural History, 1400 S. Lake Shore Dr. Chicago, IL 60605, USA.