Department of Biosciences, University of Helsinki, Helsinki, Finland.|Department of Biosciences, University of Helsinki, Helsinki, Finland.|Department of Biosciences, University of Helsinki, Helsinki, Finland.|Department of Biological and Environmental Sciences, University of Gothenburg, Gothenburg, Sweden.|Department of Biosciences, University of Helsinki, Helsinki, Finland Centre for Biodiversity Dynamics, Department of Biology, Norwegian University of Science and Technology, Trondheim, Norway.