Aix-Marseille Université, Marseille, France.|European Molecular Biology Laboratory (EMBL), Genome Biology Unit, Heidelberg, Germany.|European Molecular Biology Laboratory (EMBL), Genome Biology Unit, Heidelberg, Germany.|European Molecular Biology Laboratory (EMBL), Genome Biology Unit, Heidelberg, Germany.|Aix-Marseille Université, Marseille, France.|European Molecular Biology Laboratory (EMBL), Genome Biology Unit, Heidelberg, Germany.|Aix-Marseille Université, Marseille, France.|European Molecular Biology Laboratory (EMBL), Genome Biology Unit, Heidelberg, Germany.|Computational Systems Biology team, Institut de Biologie de lEcole Normale Supérieure (IBENS), INSERM, Ecole Normale Supérieure, PSL Research University, Paris, France.