Department of Ecology and Evolutionary Biology, University of Michigan, Ann Arbor, MI 48109, USA. benjamin.roche@ird.fr|Department of Ecology and Evolutionary Biology, University of Michigan, Ann Arbor, MI 48109, USA. benjamin.roche@ird.fr|Department of Ecology and Evolutionary Biology, University of Michigan, Ann Arbor, MI 48109, USA. benjamin.roche@ird.fr