National Institute for Mathematical and Biological Synthesis, University of Tennessee, Knoxville, Tennesse, United States of America.|Department of Forestry, Wildlife, and Fisheries, University of Tennessee, Knoxville, Tennesse, United States of America.|National Institute for Mathematical and Biological Synthesis, University of Tennessee, Knoxville, Tennesse, United States of America , Department of Microbiology, University of Tennessee, Knoxville, Tennesse, United States of America , Department of Mathematics, University of Tennessee, Knoxville, Tennesse, United States of America.