INRA, UMR1062 CBGP, F-34988 Montferrier-sur-Lez, France.|INRA, UMR1062 CBGP, F-34988 Montferrier-sur-Lez, France, IBC, F-34095 Montpellier, France.|INRA, UMR1062 CBGP, F-34988 Montferrier-sur-Lez, France.|INRA, UMR1062 CBGP, F-34988 Montferrier-sur-Lez, France.|INRA, UMR441 LIPM, F31326 Castanet Tolosan, France CNRS, UMR2594 LIPM, F31326 Castanet Tolosan, France.