Department of Biology, Washington University in St. Louis, St. Louis, MO, USA.|Department of Biomedical Engineering, Oregon Health and Science University, Portland, OR, USA.|Department of Biology, Washington University in St. Louis, St. Louis, MO, USA.|Department of Biology, Washington University in St. Louis, St. Louis, MO, USA.|Department of Biomedical Engineering, Oregon Health and Science University, Portland, OR, USA.