School of Life Sciences Research, University of Dundee, Dow Street, Dundee, DD1 5EH and Department of Computing Science, University of Glasgow, Glasgow, GL12 8QQ, UK.|School of Life Sciences Research, University of Dundee, Dow Street, Dundee, DD1 5EH and Department of Computing Science, University of Glasgow, Glasgow, GL12 8QQ, UK.|School of Life Sciences Research, University of Dundee, Dow Street, Dundee, DD1 5EH and Department of Computing Science, University of Glasgow, Glasgow, GL12 8QQ, UK.|School of Life Sciences Research, University of Dundee, Dow Street, Dundee, DD1 5EH and Department of Computing Science, University of Glasgow, Glasgow, GL12 8QQ, UK.