Joint Carnegie Mellon, University of Pittsburgh PhD Program in Computational Biology, Pittsburgh, PA, USA.|Joint Carnegie Mellon, University of Pittsburgh PhD Program in Computational Biology, Pittsburgh, PA, USA.|Joint Carnegie Mellon, University of Pittsburgh PhD Program in Computational Biology, Pittsburgh, PA, USA.|Joint Carnegie Mellon, University of Pittsburgh PhD Program in Computational Biology, Pittsburgh, PA, USA.|Joint Carnegie Mellon, University of Pittsburgh PhD Program in Computational Biology, Pittsburgh, PA, USA.