Joint Carnegie Mellon University-University of Pittsburgh Ph.D. Program in Computational Biology, Department of Biostatistics, Graduate School of Public Health, University of Pittsburgh, Pittsburgh, PA 15261, USA and Magee-Womens Research Institute, Pittsburgh, PA 15213, USA Joint Carnegie Mellon University-University of Pittsburgh Ph.D. Program in Computational Biology, Department of Biostatistics, Graduate School of Public Health, University of Pittsburgh, Pittsburgh, PA 15261, USA and Magee-Womens Research Institute, Pittsburgh, PA 15213, USA.|Joint Carnegie Mellon University-University of Pittsburgh Ph.D. Program in Computational Biology, Department of Biostatistics, Graduate School of Public Health, University of Pittsburgh, Pittsburgh, PA 15261, USA and Magee-Womens Research Institute, Pittsburgh, PA 15213, USA.|Joint Carnegie Mellon University-University of Pittsburgh Ph.D. Program in Computational Biology, Department of Biostatistics, Graduate School of Public Health, University of Pittsburgh, Pittsburgh, PA 15261, USA and Magee-Womens Research Institute, Pittsburgh, PA 15213, USA.|Joint Carnegie Mellon University-University of Pittsburgh Ph.D. Program in Computational Biology, Department of Biostatistics, Graduate School of Public Health, University of Pittsburgh, Pittsburgh, PA 15261, USA and Magee-Womens Research Institute, Pittsburgh, PA 15213, USA.|Joint Carnegie Mellon University-University of Pittsburgh Ph.D. Program in Computational Biology, Department of Biostatistics, Graduate School of Public Health, University of Pittsburgh, Pittsburgh, PA 15261, USA and Magee-Womens Research Institute, Pittsburgh, PA 15213, USA.|Joint Carnegie Mellon University-University of Pittsburgh Ph.D. Program in Computational Biology, Department of Biostatistics, Graduate School of Public Health, University of Pittsburgh, Pittsburgh, PA 15261, USA and Magee-Womens Research Institute, Pittsburgh, PA 15213, USA.|Joint Carnegie Mellon University-University of Pittsburgh Ph.D. Program in Computational Biology, Department of Biostatistics, Graduate School of Public Health, University of Pittsburgh, Pittsburgh, PA 15261, USA and Magee-Womens Research Institute, Pittsburgh, PA 15213, USA Joint Carnegie Mellon University-University of Pittsburgh Ph.D. Program in Computational Biology, Department of Biostatistics, Graduate School of Public Health, University of Pittsburgh, Pittsburgh, PA 15261, USA and Magee-Womens Research Institute, Pittsburgh, PA 15213, USA.