INRA Toulouse, Castanet Tolosan, France. matthieu.vignes@toulouse.inra.fr|INRA Toulouse, Castanet Tolosan, France. matthieu.vignes@toulouse.inra.fr|INRA Toulouse, Castanet Tolosan, France. matthieu.vignes@toulouse.inra.fr|INRA Toulouse, Castanet Tolosan, France. matthieu.vignes@toulouse.inra.fr