Department of Biochemistry, University of Wisconsin-Madison, Madison, WI 53706, USA.|Department of Biochemistry, University of Wisconsin-Madison, Madison, WI 53706, USA.|Department of Biochemistry, University of Wisconsin-Madison, Madison, WI 53706, USA.|Department of Biochemistry, University of Wisconsin-Madison, Madison, WI 53706, USA.|Department of Biochemistry, University of Wisconsin-Madison, Madison, WI 53706, USA.