Department of Anatomy and Neurobiology, Washington University, Saint Louis, Missouri, United States of America. kalpana@pcg.wustl.edu|Department of Anatomy and Neurobiology, Washington University, Saint Louis, Missouri, United States of America. kalpana@pcg.wustl.edu|Department of Anatomy and Neurobiology, Washington University, Saint Louis, Missouri, United States of America. kalpana@pcg.wustl.edu|Department of Anatomy and Neurobiology, Washington University, Saint Louis, Missouri, United States of America. kalpana@pcg.wustl.edu