Joint Carnegie Mellon - University of Pittsburgh Ph.D Program in Computational Biology, Pittsburgh, PA 15260, USA.|Joint Carnegie Mellon - University of Pittsburgh Ph.D Program in Computational Biology, Pittsburgh, PA 15260, USA.|Joint Carnegie Mellon - University of Pittsburgh Ph.D Program in Computational Biology, Pittsburgh, PA 15260, USA.|Joint Carnegie Mellon - University of Pittsburgh Ph.D Program in Computational Biology, Pittsburgh, PA 15260, USA.|Joint Carnegie Mellon - University of Pittsburgh Ph.D Program in Computational Biology, Pittsburgh, PA 15260, USA.