Götenborg University, Botanical Institute, SE-405 30, Sweden. henrik.nilsson@botany.gu.se|Götenborg University, Botanical Institute, SE-405 30, Sweden. henrik.nilsson@botany.gu.se|Götenborg University, Botanical Institute, SE-405 30, Sweden. henrik.nilsson@botany.gu.se|Götenborg University, Botanical Institute, SE-405 30, Sweden. henrik.nilsson@botany.gu.se