College of Information Science and Engineering, Hunan University, Changsha, Hunan 410082, China.|BGI, Shenzhen, Guangdong 518083, China.|College of Information Science and Engineering, Hunan University, Changsha, Hunan 410082, China.|College of Information Science and Engineering, Hunan University, Changsha, Hunan 410082, China.|BGI, Shenzhen, Guangdong 518083, China.|BGI, Shenzhen, Guangdong 518083, China.|College of Information Science and Engineering, Hunan University, Changsha, Hunan 410082, China.|College of Information Science and Engineering, Hunan University, Changsha, Hunan 410082, China.