ARK-Genomics, The Roslin Institute and Royal Dick School of Veterinary Studies, University of Edinburgh, Easter Bush, Midlothian EH25 9RG. mick.watson@roslin.ed.ac.uk|ARK-Genomics, The Roslin Institute and Royal Dick School of Veterinary Studies, University of Edinburgh, Easter Bush, Midlothian EH25 9RG. mick.watson@roslin.ed.ac.uk|ARK-Genomics, The Roslin Institute and Royal Dick School of Veterinary Studies, University of Edinburgh, Easter Bush, Midlothian EH25 9RG. mick.watson@roslin.ed.ac.uk