NASA Astrobiology Institute, Department of Biology, The Pennsylvania State University, 208 Mueller Laboratory, University Park, Pennsylvania 16802-5301, USA. jeb322@psu.edu|NASA Astrobiology Institute, Department of Biology, The Pennsylvania State University, 208 Mueller Laboratory, University Park, Pennsylvania 16802-5301, USA. jeb322@psu.edu|NASA Astrobiology Institute, Department of Biology, The Pennsylvania State University, 208 Mueller Laboratory, University Park, Pennsylvania 16802-5301, USA. jeb322@psu.edu