Department of Molecular Genetics and Microbiology, FONDAP Center for Genome Regulation, Pontificia Universidad Catolica de Chile, Santiago, Chile. tfpuelma@uc.cl|Department of Molecular Genetics and Microbiology, FONDAP Center for Genome Regulation, Pontificia Universidad Catolica de Chile, Santiago, Chile. tfpuelma@uc.cl|Department of Molecular Genetics and Microbiology, FONDAP Center for Genome Regulation, Pontificia Universidad Catolica de Chile, Santiago, Chile. tfpuelma@uc.cl