University of Tokyo, Institute of Molecular and Cellular Bioscience, Tokyo, Japan. lwy1@iam.u-tokyo.ac.jp|University of Tokyo, Institute of Molecular and Cellular Bioscience, Tokyo, Japan. lwy1@iam.u-tokyo.ac.jp|University of Tokyo, Institute of Molecular and Cellular Bioscience, Tokyo, Japan. lwy1@iam.u-tokyo.ac.jp|University of Tokyo, Institute of Molecular and Cellular Bioscience, Tokyo, Japan. lwy1@iam.u-tokyo.ac.jp