Biotechnology Program, School of Bioresources and Technology, King Mongkuts University of Technology Thonburi (Bang Khun Thian), 49 Soi Thian Thale 25, Bang Khun Thian Chai Thale Rd., Tha Kham, Bang Khun Thian, Bangkok, 10150, Thailand. marasri.rue@kmutt.ac.th.