School of Biological Sciences, University of Reading, Whiteknights, PO Box 228, Reading RG66AJ, UK. joao.lopes@reading.ac.uk|School of Biological Sciences, University of Reading, Whiteknights, PO Box 228, Reading RG66AJ, UK. joao.lopes@reading.ac.uk|School of Biological Sciences, University of Reading, Whiteknights, PO Box 228, Reading RG66AJ, UK. joao.lopes@reading.ac.uk