Department of Genetics, Development & Cell Biology, Iowa State University, Ames, Iowa 50011, USA. gaoxiang@iastate.edu|Department of Genetics, Development & Cell Biology, Iowa State University, Ames, Iowa 50011, USA. gaoxiang@iastate.edu|Department of Genetics, Development & Cell Biology, Iowa State University, Ames, Iowa 50011, USA. gaoxiang@iastate.edu|Department of Genetics, Development & Cell Biology, Iowa State University, Ames, Iowa 50011, USA. gaoxiang@iastate.edu