The Francis Crick Institute, London, United Kingdom.|Cincinnati Childrens Hospital, Division of Developmental Biology, Cincinnati, Ohio.|MRC Molecular Haematology Unit, The Weatherall Institute of Molecular Medicine, University of Oxford, John Radcliffe Hospital, Headington, Oxford.|MRC Molecular Haematology Unit, The Weatherall Institute of Molecular Medicine, University of Oxford, John Radcliffe Hospital, Headington, Oxford.|The Francis Crick Institute, London, United Kingdom.