Department of Neuroscience, Department of Physiology and Cellular Biophysics, Columbia University College of Physicians and Surgeons, New York, New York, United States of America.|Department of Neuroscience, Department of Physiology and Cellular Biophysics, Columbia University College of Physicians and Surgeons, New York, New York, United States of America.|Department of Neuroscience, Department of Physiology and Cellular Biophysics, Columbia University College of Physicians and Surgeons, New York, New York, United States of America.