MOE Key Laboratory for Biodiversity Science and Ecological Engineering and College of Life Sciences, Beijing Normal University, Beijing, China. luoyq@mail.bnu.edu.cn <luoyq@mail.bnu.edu.cn>|MOE Key Laboratory for Biodiversity Science and Ecological Engineering and College of Life Sciences, Beijing Normal University, Beijing, China. luoyq@mail.bnu.edu.cn <luoyq@mail.bnu.edu.cn>|MOE Key Laboratory for Biodiversity Science and Ecological Engineering and College of Life Sciences, Beijing Normal University, Beijing, China. luoyq@mail.bnu.edu.cn <luoyq@mail.bnu.edu.cn>|MOE Key Laboratory for Biodiversity Science and Ecological Engineering and College of Life Sciences, Beijing Normal University, Beijing, China. luoyq@mail.bnu.edu.cn <luoyq@mail.bnu.edu.cn>