Biomathematics and Statistics Scotland, University of Edinburgh Kings Buildings, Edinburgh, EH9 3JZ, Scotland, UK. mizanur@bioss.ac.uk|Biomathematics and Statistics Scotland, University of Edinburgh Kings Buildings, Edinburgh, EH9 3JZ, Scotland, UK. mizanur@bioss.ac.uk|Biomathematics and Statistics Scotland, University of Edinburgh Kings Buildings, Edinburgh, EH9 3JZ, Scotland, UK. mizanur@bioss.ac.uk