Graduate School of Agricultural and Life Sciences, University of Tokyo, Tokyo, Japan. hnishida@iu.a.u-tokyo.ac.jp|Graduate School of Agricultural and Life Sciences, University of Tokyo, Tokyo, Japan. hnishida@iu.a.u-tokyo.ac.jp|Graduate School of Agricultural and Life Sciences, University of Tokyo, Tokyo, Japan. hnishida@iu.a.u-tokyo.ac.jp|Graduate School of Agricultural and Life Sciences, University of Tokyo, Tokyo, Japan. hnishida@iu.a.u-tokyo.ac.jp|Graduate School of Agricultural and Life Sciences, University of Tokyo, Tokyo, Japan. hnishida@iu.a.u-tokyo.ac.jp