Division of Cell Biology, Department of Human Biology, Neuroscience Institute and Institute of Infectious Disease and Molecular Medicine, Faculty of Health Sciences, University of Cape Town, Cape Town, South Africa.|School of Computer Science and Applied Mathematics, University of the Witwatersrand, Johannesburg, South Africa.|Gatsby Computational Neuroscience Unit, University College London, London, United Kingdom.|Gatsby Computational Neuroscience Unit, University College London, London, United Kingdom.|Centre for Neural Circuits and Behaviour, Department of Physiology, Anatomy and Genetics, University of Oxford, Oxford, United Kingdom.|Division of Cell Biology, Department of Human Biology, Neuroscience Institute and Institute of Infectious Disease and Molecular Medicine, Faculty of Health Sciences, University of Cape Town, Cape Town, South Africa.