Biosciences, College of Life and Environmental Sciences, University of Exeter, Exeter, United Kingdom, Systems Biology Program, College of Engineering, Mathematics and Physical Sciences, University of Exeter, Exeter, United Kingdom.|Systems Biology Program, College of Engineering, Mathematics and Physical Sciences, University of Exeter, Exeter, United Kingdom.|Department of Mathematical Sciences, University of Copenhagen, Copenhagen, Denmark.|Department of Biological Sciences, Virginia Tech, Blacksburg, Virginia, United States of America.|Biosciences, College of Life and Environmental Sciences, University of Exeter, Exeter, United Kingdom.|School of Life Sciences, University of Warwick, Coventry, United Kingdom.