Department of Ecology and Evolutionary Biology, University of Arizona Tucson, P,O, Box 210088, Tucson, AZ 85721, USA. bboyle@email.arizona.edu|Department of Ecology and Evolutionary Biology, University of Arizona Tucson, P,O, Box 210088, Tucson, AZ 85721, USA. bboyle@email.arizona.edu|Department of Ecology and Evolutionary Biology, University of Arizona Tucson, P,O, Box 210088, Tucson, AZ 85721, USA. bboyle@email.arizona.edu|Department of Ecology and Evolutionary Biology, University of Arizona Tucson, P,O, Box 210088, Tucson, AZ 85721, USA. bboyle@email.arizona.edu|Department of Ecology and Evolutionary Biology, University of Arizona Tucson, P,O, Box 210088, Tucson, AZ 85721, USA. bboyle@email.arizona.edu|Department of Ecology and Evolutionary Biology, University of Arizona Tucson, P,O, Box 210088, Tucson, AZ 85721, USA. bboyle@email.arizona.edu|Department of Ecology and Evolutionary Biology, University of Arizona Tucson, P,O, Box 210088, Tucson, AZ 85721, USA. bboyle@email.arizona.edu|Department of Ecology and Evolutionary Biology, University of Arizona Tucson, P,O, Box 210088, Tucson, AZ 85721, USA. bboyle@email.arizona.edu|Department of Ecology and Evolutionary Biology, University of Arizona Tucson, P,O, Box 210088, Tucson, AZ 85721, USA. bboyle@email.arizona.edu|Department of Ecology and Evolutionary Biology, University of Arizona Tucson, P,O, Box 210088, Tucson, AZ 85721, USA. bboyle@email.arizona.edu|Department of Ecology and Evolutionary Biology, University of Arizona Tucson, P,O, Box 210088, Tucson, AZ 85721, USA. bboyle@email.arizona.edu|Department of Ecology and Evolutionary Biology, University of Arizona Tucson, P,O, Box 210088, Tucson, AZ 85721, USA. bboyle@email.arizona.edu|Department of Ecology and Evolutionary Biology, University of Arizona Tucson, P,O, Box 210088, Tucson, AZ 85721, USA. bboyle@email.arizona.edu|Department of Ecology and Evolutionary Biology, University of Arizona Tucson, P,O, Box 210088, Tucson, AZ 85721, USA. bboyle@email.arizona.edu