Department of Biostatistics Department of Genetics Department of Computer Science.|Department of Computer Science Curriculum in Bioinformatics and Computational Biology, University of North Carolina, Chapel Hill, NC 27599, USA.|College of Veterinary Medicine, Nanjing Agricultural University, Nanjing, Jiangsu 210095, China.|Department of Biostatistics Department of Genetics Department of Computer Science.|Division of Biostatistics, Department of Population Health, New York University School of Medicine, New York, NY 10016, USA.