Department of Cell & Systems Biology, University of Toronto, Toronto, Canada, Centre for the Analysis of Genome Evolution and Function, University of Toronto, Toronto, Canada.|Department of Cell & Systems Biology, University of Toronto, Toronto, Canada.|Department of Cell & Systems Biology, University of Toronto, Toronto, Canada.|Department of Cell & Systems Biology, University of Toronto, Toronto, Canada.|Département de Biologie, IBIS and PROTEO, Pavillon Charles-Eugene-Marchand, Laval University, Québec City, Canada.|Department of Molecular Biosciences, Northwestern University, Evanston, Illinois, United States of America.|Département de Biologie, IBIS and PROTEO, Pavillon Charles-Eugene-Marchand, Laval University, Québec City, Canada.|Department of Cell & Systems Biology, University of Toronto, Toronto, Canada, Centre for the Analysis of Genome Evolution and Function, University of Toronto, Toronto, Canada.