UMR 6632, Evolutionary Biology and Modeling, University of Provence, Marseille, France. philippe.gouret@univ-provence.fr|UMR 6632, Evolutionary Biology and Modeling, University of Provence, Marseille, France. philippe.gouret@univ-provence.fr|UMR 6632, Evolutionary Biology and Modeling, University of Provence, Marseille, France. philippe.gouret@univ-provence.fr