Institute of Biodiversity, Animal Health and Comparative Medicine, University of Glasgow, Glasgow, UK. anthony.ohare@stir.ac.uk.|Institute of Biodiversity, Animal Health and Comparative Medicine, University of Glasgow, Glasgow, UK. samantha.lycett@ed.ac.uk.|Institute of Biodiversity, Animal Health and Comparative Medicine, University of Glasgow, Glasgow, UK. Tom.doherty@glasgow.ac.uk.|Institute of Biodiversity, Animal Health and Comparative Medicine, University of Glasgow, Glasgow, UK. liliana.salvador@glasgow.ac.uk.|Institute of Biodiversity, Animal Health and Comparative Medicine, University of Glasgow, Glasgow, UK. rowland.kao@glasgow.ac.uk.