Computational Biology Department, Carnegie Mellon University, Pittsburgh, PA, USA.|Biomedical Engineering Department, Carnegie Mellon University, Pittsburgh, PA, USA.|Computer Science Department, Carnegie Mellon University, Pittsburgh, PA, USA.|Machine Learning Department, Carnegie Mellon University, Pittsburgh, PA, USA.|Biomedical Engineering Department, Carnegie Mellon University, Pittsburgh, PA, USA.|Division of Structural Biology, Wellcome Trust Centre for Human Genetics, University of Oxford, Oxford, UK.|Machine Learning Department, Carnegie Mellon University, Pittsburgh, PA, USA.