School of Biological Sciences, University of Reading, UK. R.A.Vos@reading.ac.uk|School of Biological Sciences, University of Reading, UK. R.A.Vos@reading.ac.uk|School of Biological Sciences, University of Reading, UK. R.A.Vos@reading.ac.uk|School of Biological Sciences, University of Reading, UK. R.A.Vos@reading.ac.uk|School of Biological Sciences, University of Reading, UK. R.A.Vos@reading.ac.uk