School of Physiology, Pharmacology and Neuroscience, University of Bristol, Bristol, United Kingdom.|Wellcome Trust Centre for Neuroimaging, UCL, London, United Kingdom.|School of Physiology, Pharmacology and Neuroscience, University of Bristol, Bristol, United Kingdom.|School of Physiology, Pharmacology and Neuroscience, University of Bristol, Bristol, United Kingdom.|School of Physiology, Pharmacology and Neuroscience, University of Bristol, Bristol, United Kingdom.