Department of Integrative Biology, Department of Statistics, University of California, Berkeley, CA 94720, USA, Department of Evolution and Ecology, University of California, Davis, CA 95616, USA and Department of Mathematics, Stockholm University, Stockholm, SE-106 91, Sweden.|Department of Evolution and Ecology, University of California, Davis, CA 95616, USA and.|Department of Evolution and Ecology, University of California, Davis, CA 95616, USA and.