Centre for Plant Integrative Biology, University of Nottingham, Sutton Bonington Campus, Nottingham, UK.|Centre for Plant Integrative Biology, University of Nottingham, Sutton Bonington Campus, Nottingham, UK.|Centre for Plant Integrative Biology, University of Nottingham, Sutton Bonington Campus, Nottingham, UK.|Centre for Plant Integrative Biology, University of Nottingham, Sutton Bonington Campus, Nottingham, UK.