John Curtin School of Medical Research, The Australian National University, Canberra 0200 ACT, Australia. matthew.wakefield@anu.edu.au|John Curtin School of Medical Research, The Australian National University, Canberra 0200 ACT, Australia. matthew.wakefield@anu.edu.au|John Curtin School of Medical Research, The Australian National University, Canberra 0200 ACT, Australia. matthew.wakefield@anu.edu.au