Department of Plant Biology, University of Georgia, Athens, GA 30602, USA. chapmanb@uga.edu|Department of Plant Biology, University of Georgia, Athens, GA 30602, USA. chapmanb@uga.edu|Department of Plant Biology, University of Georgia, Athens, GA 30602, USA. chapmanb@uga.edu|Department of Plant Biology, University of Georgia, Athens, GA 30602, USA. chapmanb@uga.edu