Department of Biological Sciences, University of South Carolina Columbia, SC 29208, USA. austin@biol.sc.edu|Department of Biological Sciences, University of South Carolina Columbia, SC 29208, USA. austin@biol.sc.edu|Department of Biological Sciences, University of South Carolina Columbia, SC 29208, USA. austin@biol.sc.edu|Department of Biological Sciences, University of South Carolina Columbia, SC 29208, USA. austin@biol.sc.edu|Department of Biological Sciences, University of South Carolina Columbia, SC 29208, USA. austin@biol.sc.edu