Biomathematics & Statistics Scotland, SCRI, Dundee, DD2 5DA, UK. topali@bioss.ac.uk|Biomathematics & Statistics Scotland, SCRI, Dundee, DD2 5DA, UK. topali@bioss.ac.uk|Biomathematics & Statistics Scotland, SCRI, Dundee, DD2 5DA, UK. topali@bioss.ac.uk|Biomathematics & Statistics Scotland, SCRI, Dundee, DD2 5DA, UK. topali@bioss.ac.uk|Biomathematics & Statistics Scotland, SCRI, Dundee, DD2 5DA, UK. topali@bioss.ac.uk|Biomathematics & Statistics Scotland, SCRI, Dundee, DD2 5DA, UK. topali@bioss.ac.uk