School of Biological Sciences, Ashworth Laboratories, Kings Buildings, West Mains Rd, University of Edinburgh, Edinburgh EH9 3JT, UK. jparkin@sickkids.ca|School of Biological Sciences, Ashworth Laboratories, Kings Buildings, West Mains Rd, University of Edinburgh, Edinburgh EH9 3JT, UK. jparkin@sickkids.ca|School of Biological Sciences, Ashworth Laboratories, Kings Buildings, West Mains Rd, University of Edinburgh, Edinburgh EH9 3JT, UK. jparkin@sickkids.ca|School of Biological Sciences, Ashworth Laboratories, Kings Buildings, West Mains Rd, University of Edinburgh, Edinburgh EH9 3JT, UK. jparkin@sickkids.ca|School of Biological Sciences, Ashworth Laboratories, Kings Buildings, West Mains Rd, University of Edinburgh, Edinburgh EH9 3JT, UK. jparkin@sickkids.ca|School of Biological Sciences, Ashworth Laboratories, Kings Buildings, West Mains Rd, University of Edinburgh, Edinburgh EH9 3JT, UK. jparkin@sickkids.ca