Evolution and Ecology Program, International Institute for Applied Systems Analysis (IIASA), Laxenburg, Austria. boza@iiasa.ac.at|Evolution and Ecology Program, International Institute for Applied Systems Analysis (IIASA), Laxenburg, Austria. boza@iiasa.ac.at|Evolution and Ecology Program, International Institute for Applied Systems Analysis (IIASA), Laxenburg, Austria. boza@iiasa.ac.at|Evolution and Ecology Program, International Institute for Applied Systems Analysis (IIASA), Laxenburg, Austria. boza@iiasa.ac.at