Laboratory of Genetics, University of Wisconsin-Madison, Madison, WI, United States of America.|Department of Biomolecular Chemistry, University of Wisconsin-Madison, Madison, WI, United States of America.|Wisconsin Institute for Discovery, University of Wisconsin-Madison, Madison, WI, United States of America.|Great Lakes Bioenergy Research Center, University of Wisconsin-Madison, Madison, WI, United States of America.|Department of Biostatistics and Medical Informatics, University of Wisconsin -Madison, Madison, WI, United States of America.|Department of Biomolecular Chemistry, University of Wisconsin-Madison, Madison, WI, United States of America.|Laboratory of Genetics, University of Wisconsin-Madison, Madison, WI, United States of America.