Department of Physiology and the Center for Integrative Neuroscience, University of California San Francisco, San Francisco, California, USA. makin@phy.ucsf.edu|Department of Physiology and the Center for Integrative Neuroscience, University of California San Francisco, San Francisco, California, USA. makin@phy.ucsf.edu|Department of Physiology and the Center for Integrative Neuroscience, University of California San Francisco, San Francisco, California, USA. makin@phy.ucsf.edu