Courant Institute of Mathematical Sciences, New York University, New York, New York, United States of America.|Program in Developmental Biology, Sloan Kettering Institute, Memorial Sloan Kettering Cancer, New York, New York, United States of America.|Program in Developmental Biology, Sloan Kettering Institute, Memorial Sloan Kettering Cancer, New York, New York, United States of America.|Courant Institute of Mathematical Sciences, New York University, New York, New York, United States of America.