Department of Ecology and Evolution, University of Chicago, Chicago, IL, United States of America.|Department of Ecology and Evolution, University of Chicago, Chicago, IL, United States of America.|School of Biosciences, Bio21 Institute, The University of Melbourne, Melbourne, AU.|School of Biosciences, Bio21 Institute, The University of Melbourne, Melbourne, AU.|Department of Ecology and Evolution, University of Chicago, Chicago, IL, United States of America.