Department of Microbiology and Biochemistry, University of Victoria, Victoria, BC, V8W 3P6, Canada. jwhitney@gmail.com|Department of Microbiology and Biochemistry, University of Victoria, Victoria, BC, V8W 3P6, Canada. jwhitney@gmail.com|Department of Microbiology and Biochemistry, University of Victoria, Victoria, BC, V8W 3P6, Canada. jwhitney@gmail.com