Evolutionary Genetics, Free University of Brussels (ULB), cp 300, Institute of Molecular Biology and Medicine, rue Jeener & Brachet 12, B-6041 Gosselies, Belgium. apl@dbm.ulb.ac.be|Evolutionary Genetics, Free University of Brussels (ULB), cp 300, Institute of Molecular Biology and Medicine, rue Jeener & Brachet 12, B-6041 Gosselies, Belgium. apl@dbm.ulb.ac.be