Graduate School of Biostudies, Kyoto University, Kyoto, Japan. ksugimura@brain.riken.jp|Graduate School of Biostudies, Kyoto University, Kyoto, Japan. ksugimura@brain.riken.jp|Graduate School of Biostudies, Kyoto University, Kyoto, Japan. ksugimura@brain.riken.jp|Graduate School of Biostudies, Kyoto University, Kyoto, Japan. ksugimura@brain.riken.jp