Department of Molecular and Cell Biology, University of California Berkeley, Berkeley, California, USA. myandell@genetics.utah|Department of Molecular and Cell Biology, University of California Berkeley, Berkeley, California, USA. myandell@genetics.utah|Department of Molecular and Cell Biology, University of California Berkeley, Berkeley, California, USA. myandell@genetics.utah|Department of Molecular and Cell Biology, University of California Berkeley, Berkeley, California, USA. myandell@genetics.utah|Department of Molecular and Cell Biology, University of California Berkeley, Berkeley, California, USA. myandell@genetics.utah|Department of Molecular and Cell Biology, University of California Berkeley, Berkeley, California, USA. myandell@genetics.utah|Department of Molecular and Cell Biology, University of California Berkeley, Berkeley, California, USA. myandell@genetics.utah|Department of Molecular and Cell Biology, University of California Berkeley, Berkeley, California, USA. myandell@genetics.utah