Evolutionary Theory Group, Max-Planck-Institute for Evolutionary Biology, Plön, Germany. altrock@evolbio.mpg.de|Evolutionary Theory Group, Max-Planck-Institute for Evolutionary Biology, Plön, Germany. altrock@evolbio.mpg.de|Evolutionary Theory Group, Max-Planck-Institute for Evolutionary Biology, Plön, Germany. altrock@evolbio.mpg.de