Laboratory of Theoretical Biology, Research Center for Dynamic Living Systems, Graduate School of Biostudies, Kyoto University, Yoshidakonoecho, Sakyo, Kyoto, Japan.|Gene Regulation Research, Division of Biological Science, Nara Institute of Science and Technology, Takayama, Nara, Japan.|Gene Regulation Research, Division of Biological Science, Nara Institute of Science and Technology, Takayama, Nara, Japan.|Graduate School of Informatics, Kyoto University, Yoshidahonmachi, Sakyo, Kyoto, Japan.|Gene Regulation Research, Division of Biological Science, Nara Institute of Science and Technology, Takayama, Nara, Japan.|Gene Regulation Research, Division of Biological Science, Nara Institute of Science and Technology, Takayama, Nara, Japan.