School of Computer Science and Shanghai Key Lab of Intelligent Information Processing, Fudan University, Shanghai 200433, China.|School of Computer Science and Shanghai Key Lab of Intelligent Information Processing, Fudan University, Shanghai 200433, China.|Department of Computer Science, University of Virginia, Charlottesville 22904-4740, USA.|Department of Computer Science, University of Illinois at Urbana-Champaign, IL 61801, USA.|Bioinformatics Center, Kyoto University, Institute for Chemical Research, Uji 611-0011, Japan Department of Computer Science, Aalto University, Finland.|School of Computer Science and Shanghai Key Lab of Intelligent Information Processing, Fudan University, Shanghai 200433, China Centre for Computational System Biology, Fudan University, Shanghai 200433, China.