Department of Ecology and Evolutionary Biology, University of Michigan, Ann Arbor, Michigan, USA.|Department of Ecology and Evolutionary Biology, University of Michigan, Ann Arbor, Michigan, USA.|Department of Ecology and Evolutionary Biology, University of Michigan, Ann Arbor, Michigan, USA.|Department of Ecology and Evolutionary Biology, University of Michigan, Ann Arbor, Michigan, USA.|Department of Ecology and Evolutionary Biology, University of Michigan, Ann Arbor, Michigan, USA.|Department of Ecology and Evolutionary Biology, University of Michigan, Ann Arbor, Michigan, USA.|Department of Ecology and Evolutionary Biology, University of Michigan, Ann Arbor, Michigan, USA.