Ecosystems and Environment Research Centre, School of Environment and Life Sciences, University of Salford, Greater Manchester, United Kingdom.|Computational and Systems Biology, Rothamsted Research, Harpenden, Hertfordshire, United Kingdom.|USDA Agricultural Research Service, Fort Pierce, Florida, United States of America.|Computational and Systems Biology, Rothamsted Research, Harpenden, Hertfordshire, United Kingdom.|Ecosystems and Environment Research Centre, School of Environment and Life Sciences, University of Salford, Greater Manchester, United Kingdom.