Department of Biology, James Madison University, 820 Madison Dr, MSC 7801, Harrisonburg, VA 22807 USA. cresawsg@jmu.edu|Department of Biology, James Madison University, 820 Madison Dr, MSC 7801, Harrisonburg, VA 22807 USA. cresawsg@jmu.edu|Department of Biology, James Madison University, 820 Madison Dr, MSC 7801, Harrisonburg, VA 22807 USA. cresawsg@jmu.edu|Department of Biology, James Madison University, 820 Madison Dr, MSC 7801, Harrisonburg, VA 22807 USA. cresawsg@jmu.edu|Department of Biology, James Madison University, 820 Madison Dr, MSC 7801, Harrisonburg, VA 22807 USA. cresawsg@jmu.edu|Department of Biology, James Madison University, 820 Madison Dr, MSC 7801, Harrisonburg, VA 22807 USA. cresawsg@jmu.edu