Department of Ecology and Evolutionary Biology, Princeton University, Princeton, New Jersey, United States of America. simon.garnier@gmail.com|Department of Ecology and Evolutionary Biology, Princeton University, Princeton, New Jersey, United States of America. simon.garnier@gmail.com|Department of Ecology and Evolutionary Biology, Princeton University, Princeton, New Jersey, United States of America. simon.garnier@gmail.com|Department of Ecology and Evolutionary Biology, Princeton University, Princeton, New Jersey, United States of America. simon.garnier@gmail.com|Department of Ecology and Evolutionary Biology, Princeton University, Princeton, New Jersey, United States of America. simon.garnier@gmail.com|Department of Ecology and Evolutionary Biology, Princeton University, Princeton, New Jersey, United States of America. simon.garnier@gmail.com