Arnold Arboretum, Harvard University, 22 Divinity Avenue, Cambridge, MA 02138, USA. cwebb@oeb.harvard.edu|Arnold Arboretum, Harvard University, 22 Divinity Avenue, Cambridge, MA 02138, USA. cwebb@oeb.harvard.edu|Arnold Arboretum, Harvard University, 22 Divinity Avenue, Cambridge, MA 02138, USA. cwebb@oeb.harvard.edu