Department of Biology, Boston College, Chestnut Hill, MA 02467, USA.|Department of Biology, Boston College, Chestnut Hill, MA 02467, USA.|Department of Biology, Boston College, Chestnut Hill, MA 02467, USA.|Department of Biology, Boston College, Chestnut Hill, MA 02467, USA.|Department of Biology, Boston College, Chestnut Hill, MA 02467, USA.