Hunan Provincial Key Laboratory for Biology and Control of Plant Diseases and Insect Pests, Changsha, China. chenyuan0510@126.com.|Biotechnology Research Center, Hunan Academy of Agricultural Sciences, Changsha, China. wanglifeng112358@163.com.|Hunan Provincial Key Laboratory for Germplasm Innovation and Utilization of Crop, Hunan Agricultural University, Changsha, China. lancy0829@163.com.|Hunan Provincial Key Laboratory for Germplasm Innovation and Utilization of Crop, Hunan Agricultural University, Changsha, China. hongyan_zhang6@aliyun.com.|Hunan Provincial Key Laboratory for Biology and Control of Plant Diseases and Insect Pests, Changsha, China. zhmyuan@sina.com.