Institute for Neural Computation, University of California at San Diego, La Jolla, California, United States of America.|Institute for Neural Computation, University of California at San Diego, La Jolla, California, United States of America.|Institute for Neural Computation, University of California at San Diego, La Jolla, California, United States of America, Graduate Program in Neurosciences, University of California at San Diego, La Jolla, California, United States of America.|Systems Neurobiology Laboratories, Salk Institute for Biological Studies, La Jolla, California, United States of America.