Department of Ecology and Evolutionary Biology, Yale University, New Haven, CT 06520-8106, USA. junhyong.kim@yale.edu|Department of Ecology and Evolutionary Biology, Yale University, New Haven, CT 06520-8106, USA. junhyong.kim@yale.edu|Department of Ecology and Evolutionary Biology, Yale University, New Haven, CT 06520-8106, USA. junhyong.kim@yale.edu|Department of Ecology and Evolutionary Biology, Yale University, New Haven, CT 06520-8106, USA. junhyong.kim@yale.edu|Department of Ecology and Evolutionary Biology, Yale University, New Haven, CT 06520-8106, USA. junhyong.kim@yale.edu