Research School of Biological Sciences, Australian National University, PO Box 475, Canberra ACT 2601, Australia. andrew.james@anu.edu.au|Research School of Biological Sciences, Australian National University, PO Box 475, Canberra ACT 2601, Australia. andrew.james@anu.edu.au|Research School of Biological Sciences, Australian National University, PO Box 475, Canberra ACT 2601, Australia. andrew.james@anu.edu.au|Research School of Biological Sciences, Australian National University, PO Box 475, Canberra ACT 2601, Australia. andrew.james@anu.edu.au