Department of Life Sciences, Nottingham Trent University, Clifton Lane, Clifton, Nottingham NG11 8NS, UK. graham.balls@ntu.ac.uk|Department of Life Sciences, Nottingham Trent University, Clifton Lane, Clifton, Nottingham NG11 8NS, UK. graham.balls@ntu.ac.uk|Department of Life Sciences, Nottingham Trent University, Clifton Lane, Clifton, Nottingham NG11 8NS, UK. graham.balls@ntu.ac.uk|Department of Life Sciences, Nottingham Trent University, Clifton Lane, Clifton, Nottingham NG11 8NS, UK. graham.balls@ntu.ac.uk|Department of Life Sciences, Nottingham Trent University, Clifton Lane, Clifton, Nottingham NG11 8NS, UK. graham.balls@ntu.ac.uk|Department of Life Sciences, Nottingham Trent University, Clifton Lane, Clifton, Nottingham NG11 8NS, UK. graham.balls@ntu.ac.uk|Department of Life Sciences, Nottingham Trent University, Clifton Lane, Clifton, Nottingham NG11 8NS, UK. graham.balls@ntu.ac.uk|Department of Life Sciences, Nottingham Trent University, Clifton Lane, Clifton, Nottingham NG11 8NS, UK. graham.balls@ntu.ac.uk|Department of Life Sciences, Nottingham Trent University, Clifton Lane, Clifton, Nottingham NG11 8NS, UK. graham.balls@ntu.ac.uk|Department of Life Sciences, Nottingham Trent University, Clifton Lane, Clifton, Nottingham NG11 8NS, UK. graham.balls@ntu.ac.uk|Department of Life Sciences, Nottingham Trent University, Clifton Lane, Clifton, Nottingham NG11 8NS, UK. graham.balls@ntu.ac.uk