Department of Neurobiology and Physiology, Northwestern University, Evanston, Illinois, United States of America.|Department of Neurobiology and Physiology, Northwestern University, Evanston, Illinois, United States of America.|Department of Neurobiology and Physiology, Northwestern University, Evanston, Illinois, United States of America.|Department of Neurobiology and Physiology, Northwestern University, Evanston, Illinois, United States of America.