Institute for Cell and Molecular Biosciences, Newcastle University, Newcastle, UK.|Department of Biosciences, Durham University, Durham, UK.|Institute of Cellular Medicine, Newcastle University, Newcastle, UK.|Institute for Cell and Molecular Biosciences, Newcastle University, Newcastle, UK.|The Proctor & Gamble Company, Cincinnati, USA.|The Proctor & Gamble Company, Cincinnati, USA.|The Proctor & Gamble Company, Cincinnati, USA.|Department of Biosciences, Durham University, Durham, UK.|Institute for Cell and Molecular Biosciences, Newcastle University, Newcastle, UK.