Department of Genetics, University of Cambridge, Cambridge, UK. rh497@cam.ac.uk|Department of Genetics, University of Cambridge, Cambridge, UK. rh497@cam.ac.uk|Department of Genetics, University of Cambridge, Cambridge, UK. rh497@cam.ac.uk|Department of Genetics, University of Cambridge, Cambridge, UK. rh497@cam.ac.uk|Department of Genetics, University of Cambridge, Cambridge, UK. rh497@cam.ac.uk|Department of Genetics, University of Cambridge, Cambridge, UK. rh497@cam.ac.uk|Department of Genetics, University of Cambridge, Cambridge, UK. rh497@cam.ac.uk