Department of Veterinary Microbiology and Preventive Medicine, Iowa State University, Ames, Iowa, USA. gregory@iastate.edu.|Department of Veterinary Microbiology and Preventive Medicine, Iowa State University, Ames, Iowa, USA. gregory@iastate.edu.|Department of Veterinary Microbiology and Preventive Medicine, Iowa State University, Ames, Iowa, USA. gregory@iastate.edu.|Department of Veterinary Microbiology and Preventive Medicine, Iowa State University, Ames, Iowa, USA. gregory@iastate.edu.