School of Biology and Environmental Science, University College Dublin, Belfield, Dublin 4, Ireland. mbekaert@gmail.com|School of Biology and Environmental Science, University College Dublin, Belfield, Dublin 4, Ireland. mbekaert@gmail.com|School of Biology and Environmental Science, University College Dublin, Belfield, Dublin 4, Ireland. mbekaert@gmail.com|School of Biology and Environmental Science, University College Dublin, Belfield, Dublin 4, Ireland. mbekaert@gmail.com