Department of Biology and Institute for CyberScience, Pennsylvania State University, University Park, PA, USA.|Department of Ecology and Evolutionary Biology, University of California, Los Angeles, CA, USA.|Department of Biological Statistics and Computational Biology, Cornell University, Ithaca, NY, USA.|Department Biologie II, Ludwig-Maximilians-Universität München, Planegg-Martinsried, Germany and.|Department of Integrative Biology, University of California, Berkeley, CA, USA.