Wellcome Trust Centre for Cell Biology and School of Biological Sciences, The University of Edinburgh, Edinburgh EH9 3JR, UK. jan.wildenhain@ed.ac.uk|Wellcome Trust Centre for Cell Biology and School of Biological Sciences, The University of Edinburgh, Edinburgh EH9 3JR, UK. jan.wildenhain@ed.ac.uk|Wellcome Trust Centre for Cell Biology and School of Biological Sciences, The University of Edinburgh, Edinburgh EH9 3JR, UK. jan.wildenhain@ed.ac.uk