Department of Biology, The University of Texas at Arlington, Arlington, TX 76019, USA.|Department of Biology, The University of Texas at Arlington, Arlington, TX 76019, USA.|Department of Biology, The University of Texas at Arlington, Arlington, TX 76019, USA.|Department of Ecology, Evolution & Behavior, University of Minnesota, Saint Paul, MN 55108, USA.|Department of Biology, The University of Texas at Arlington, Arlington, TX 76019, USA.