Department of Biosciences, Department of Neurobiology, A.I.Virtanen Institute for Molecular Sciences, Biocenter Finland, Kuopio, Finland. liisa.heikkinen@uef.fi|Department of Biosciences, Department of Neurobiology, A.I.Virtanen Institute for Molecular Sciences, Biocenter Finland, Kuopio, Finland. liisa.heikkinen@uef.fi|Department of Biosciences, Department of Neurobiology, A.I.Virtanen Institute for Molecular Sciences, Biocenter Finland, Kuopio, Finland. liisa.heikkinen@uef.fi