Department of Zoology and Ecology, University of Navarra, E-31080 Pamplona, Spain. artarip@unav.es|Department of Zoology and Ecology, University of Navarra, E-31080 Pamplona, Spain. artarip@unav.es|Department of Zoology and Ecology, University of Navarra, E-31080 Pamplona, Spain. artarip@unav.es