Joint CMU-Pitt PhD Program in Computational Biology, Lane Center for Computational Biology, School of Computer Science, Language Technologies Institute and Department of Machine Learning, Carnegie Mellon University, Pittsburgh, PA 15213, USA.|Joint CMU-Pitt PhD Program in Computational Biology, Lane Center for Computational Biology, School of Computer Science, Language Technologies Institute and Department of Machine Learning, Carnegie Mellon University, Pittsburgh, PA 15213, USA.|Joint CMU-Pitt PhD Program in Computational Biology, Lane Center for Computational Biology, School of Computer Science, Language Technologies Institute and Department of Machine Learning, Carnegie Mellon University, Pittsburgh, PA 15213, USA.|Joint CMU-Pitt PhD Program in Computational Biology, Lane Center for Computational Biology, School of Computer Science, Language Technologies Institute and Department of Machine Learning, Carnegie Mellon University, Pittsburgh, PA 15213, USA.|Joint CMU-Pitt PhD Program in Computational Biology, Lane Center for Computational Biology, School of Computer Science, Language Technologies Institute and Department of Machine Learning, Carnegie Mellon University, Pittsburgh, PA 15213, USA.