Univ. Grenoble Alpes, TIMC-IMAG, Grenoble, F-38000, France. duforetn@berkeley.edu.|Department of Evolutionary Biology, Evolutionary Biology Centre, Uppsala University, Uppsala, Sweden. lucie.gattepaille@gmail.com.|Univ. Grenoble Alpes, TIMC-IMAG, Grenoble, F-38000, France. michael.blum@imag.fr.|Department of Evolutionary Biology, Evolutionary Biology Centre, Uppsala University, Uppsala, Sweden. mattias.jakobsson@ebc.uu.se.