Department of Mathematics, University of California Los Angeles, Los Angeles, California, United States of America.|Department of Mathematics, University of California Los Angeles, Los Angeles, California, United States of America.|Department of Bioengineering, School of Engineering & Applied Science, University of California Los Angeles, Los Angeles, California, United States of America.|Department of Bioengineering, School of Engineering & Applied Science, University of California Los Angeles, Los Angeles, California, United States of America.|Department of Life Science, National Taiwan University, Taipei, Taiwan, Republic of China.|Department of Biomathematics, University of California Los Angeles, Los Angeles, California, United States of America.|Institute of Cellular and Organismic Biology, Academia Sinica, Nankang, Taipei, Taiwan, Republic of China.|Department of Bioengineering, School of Engineering & Applied Science, University of California Los Angeles, Los Angeles, California, United States of America.|Department of Mathematics, University of California Los Angeles, Los Angeles, California, United States of America.