Department of Gene Regulation, Stem Cells, and Cancer, Centre for Genomic Regulation (CRG) and Department of Experimental and Health Sciences (CEXS), Universitat Pompeu Fabra, 08003 Barcelona, SpainDepartment of Gene Regulation, Stem Cells, and Cancer, Centre for Genomic Regulation (CRG) and Department of Experimental and Health Sciences (CEXS), Universitat Pompeu Fabra, 08003 Barcelona, Spain.