School of Biological Sciences, University of Bristol, Bristol, BS8 1UG, UK. Paul.Wilkinson@bristol.ac.uk|School of Biological Sciences, University of Bristol, Bristol, BS8 1UG, UK. Paul.Wilkinson@bristol.ac.uk|School of Biological Sciences, University of Bristol, Bristol, BS8 1UG, UK. Paul.Wilkinson@bristol.ac.uk|School of Biological Sciences, University of Bristol, Bristol, BS8 1UG, UK. Paul.Wilkinson@bristol.ac.uk|School of Biological Sciences, University of Bristol, Bristol, BS8 1UG, UK. Paul.Wilkinson@bristol.ac.uk|School of Biological Sciences, University of Bristol, Bristol, BS8 1UG, UK. Paul.Wilkinson@bristol.ac.uk|School of Biological Sciences, University of Bristol, Bristol, BS8 1UG, UK. Paul.Wilkinson@bristol.ac.uk