Research Centre for Biodiversity, Academia Sinica, Taipei, Taiwan. cckenneth@gmail.com|Research Centre for Biodiversity, Academia Sinica, Taipei, Taiwan. cckenneth@gmail.com|Research Centre for Biodiversity, Academia Sinica, Taipei, Taiwan. cckenneth@gmail.com|Research Centre for Biodiversity, Academia Sinica, Taipei, Taiwan. cckenneth@gmail.com