Department of Biology, University of Nevada, Reno, Nevada, USA. hoelzer@unr.edu|Department of Biology, University of Nevada, Reno, Nevada, USA. hoelzer@unr.edu|Department of Biology, University of Nevada, Reno, Nevada, USA. hoelzer@unr.edu|Department of Biology, University of Nevada, Reno, Nevada, USA. hoelzer@unr.edu