Sir Harold Mitchell Building, School of Biology, University of St Andrews, St Andrews, Fife KY16 9TH, UK. db60@st-andrews.ac.uk|Sir Harold Mitchell Building, School of Biology, University of St Andrews, St Andrews, Fife KY16 9TH, UK. db60@st-andrews.ac.uk|Sir Harold Mitchell Building, School of Biology, University of St Andrews, St Andrews, Fife KY16 9TH, UK. db60@st-andrews.ac.uk|Sir Harold Mitchell Building, School of Biology, University of St Andrews, St Andrews, Fife KY16 9TH, UK. db60@st-andrews.ac.uk|Sir Harold Mitchell Building, School of Biology, University of St Andrews, St Andrews, Fife KY16 9TH, UK. db60@st-andrews.ac.uk|Sir Harold Mitchell Building, School of Biology, University of St Andrews, St Andrews, Fife KY16 9TH, UK. db60@st-andrews.ac.uk|Sir Harold Mitchell Building, School of Biology, University of St Andrews, St Andrews, Fife KY16 9TH, UK. db60@st-andrews.ac.uk