Division of Biology, California Institute of Technology, Pasadena, CA 91125, USA.|Division of Biology, California Institute of Technology, Pasadena, CA 91125, USA.|Division of Biology, California Institute of Technology, Pasadena, CA 91125, USA.|Division of Biology, California Institute of Technology, Pasadena, CA 91125, USA.|Division of Biology, California Institute of Technology, Pasadena, CA 91125, USA.