Sorbonne Universités, UPMC Univ Paris 06, CNRS, Institute of Intelligent Systems and Robotics (ISIR), Paris, France.|Institut des Sciences de lEvolution, Université de Montpellier, CNRS, IRD, EPHE, CC065, Pl. E. Bataillon, Montpellier, France.|Sorbonne Universités, UPMC Univ Paris 06, CNRS, Institute of Intelligent Systems and Robotics (ISIR), Paris, France.