Department of Genetics, Development and Cell Biology, Iowa State University, 2112 Molecular Biology Building, Ames, IA 50011-3260, USA. vbrendel@iastate.edu|Department of Genetics, Development and Cell Biology, Iowa State University, 2112 Molecular Biology Building, Ames, IA 50011-3260, USA. vbrendel@iastate.edu|Department of Genetics, Development and Cell Biology, Iowa State University, 2112 Molecular Biology Building, Ames, IA 50011-3260, USA. vbrendel@iastate.edu