Division of Biology and Biological Engineering, California Institute of Technology, Pasadena, California, United States of America.|Division of Biology and Biological Engineering, California Institute of Technology, Pasadena, California, United States of America, Sobell Department of Motor Neuroscience and Movement Disorders, University College London, London, United Kingdom.|Division of Biology and Biological Engineering, California Institute of Technology, Pasadena, California, United States of America.