Crick-Jacobs Center for Theoretical and Computational Biology, Salk Institute, La Jolla, California, USA. geneyeo@salk.edu|Crick-Jacobs Center for Theoretical and Computational Biology, Salk Institute, La Jolla, California, USA. geneyeo@salk.edu|Crick-Jacobs Center for Theoretical and Computational Biology, Salk Institute, La Jolla, California, USA. geneyeo@salk.edu|Crick-Jacobs Center for Theoretical and Computational Biology, Salk Institute, La Jolla, California, USA. geneyeo@salk.edu|Crick-Jacobs Center for Theoretical and Computational Biology, Salk Institute, La Jolla, California, USA. geneyeo@salk.edu|Crick-Jacobs Center for Theoretical and Computational Biology, Salk Institute, La Jolla, California, USA. geneyeo@salk.edu|Crick-Jacobs Center for Theoretical and Computational Biology, Salk Institute, La Jolla, California, USA. geneyeo@salk.edu