School of Biosciences, Hatherly building, Exeter EX4 4PS, University of Exeter, UK. z.r.yang@ex.ac.uk|School of Biosciences, Hatherly building, Exeter EX4 4PS, University of Exeter, UK. z.r.yang@ex.ac.uk|School of Biosciences, Hatherly building, Exeter EX4 4PS, University of Exeter, UK. z.r.yang@ex.ac.uk|School of Biosciences, Hatherly building, Exeter EX4 4PS, University of Exeter, UK. z.r.yang@ex.ac.uk|School of Biosciences, Hatherly building, Exeter EX4 4PS, University of Exeter, UK. z.r.yang@ex.ac.uk