National Center for Ecological Analysis and Synthesis, University of California, Santa Barbara, Santa Barbara, California, United States of America. melian@nceas.ucsb.edu|National Center for Ecological Analysis and Synthesis, University of California, Santa Barbara, Santa Barbara, California, United States of America. melian@nceas.ucsb.edu|National Center for Ecological Analysis and Synthesis, University of California, Santa Barbara, Santa Barbara, California, United States of America. melian@nceas.ucsb.edu|National Center for Ecological Analysis and Synthesis, University of California, Santa Barbara, Santa Barbara, California, United States of America. melian@nceas.ucsb.edu|National Center for Ecological Analysis and Synthesis, University of California, Santa Barbara, Santa Barbara, California, United States of America. melian@nceas.ucsb.edu