Computational Biology Department, Carnegie Mellon University, Pittsburgh, Pennsylvania, United States of America.|Center for Bioimage Informatics, Carnegie Mellon University, Pittsburgh, Pennsylvania, United States of America.|Computational Biology Department, Carnegie Mellon University, Pittsburgh, Pennsylvania, United States of America.|Computational Biology Department, Carnegie Mellon University, Pittsburgh, Pennsylvania, United States of America.|Computational Biology Department, Carnegie Mellon University, Pittsburgh, Pennsylvania, United States of America.