Image Bioinformatics Research Group, Department of Zoology, University of Oxford, The Tinbergen Building, South Parks Road, Oxford, OX1 3PS, UK. jun.zhao@zoo.ox.ac.uk|Image Bioinformatics Research Group, Department of Zoology, University of Oxford, The Tinbergen Building, South Parks Road, Oxford, OX1 3PS, UK. jun.zhao@zoo.ox.ac.uk|Image Bioinformatics Research Group, Department of Zoology, University of Oxford, The Tinbergen Building, South Parks Road, Oxford, OX1 3PS, UK. jun.zhao@zoo.ox.ac.uk|Image Bioinformatics Research Group, Department of Zoology, University of Oxford, The Tinbergen Building, South Parks Road, Oxford, OX1 3PS, UK. jun.zhao@zoo.ox.ac.uk