Department of Molecular, Cellular & Developmental Biology, University of Michigan, Ann Arbor, MI 48109-1048, USA. catqliu@umich.edu|Department of Molecular, Cellular & Developmental Biology, University of Michigan, Ann Arbor, MI 48109-1048, USA. catqliu@umich.edu|Department of Molecular, Cellular & Developmental Biology, University of Michigan, Ann Arbor, MI 48109-1048, USA. catqliu@umich.edu|Department of Molecular, Cellular & Developmental Biology, University of Michigan, Ann Arbor, MI 48109-1048, USA. catqliu@umich.edu