SUPA, Institute for Complex Systems and Mathematical Biology, Kings College, University of Aberdeen, Aberdeen, United Kingdom. l.ciandrini@abdn.ac.uk|SUPA, Institute for Complex Systems and Mathematical Biology, Kings College, University of Aberdeen, Aberdeen, United Kingdom. l.ciandrini@abdn.ac.uk|SUPA, Institute for Complex Systems and Mathematical Biology, Kings College, University of Aberdeen, Aberdeen, United Kingdom. l.ciandrini@abdn.ac.uk