Department of Ecology and Evolutionary Biology, Princeton University, Princeton, New Jersey, United States of America. tunstrom@princeton.edu|Department of Ecology and Evolutionary Biology, Princeton University, Princeton, New Jersey, United States of America. tunstrom@princeton.edu|Department of Ecology and Evolutionary Biology, Princeton University, Princeton, New Jersey, United States of America. tunstrom@princeton.edu|Department of Ecology and Evolutionary Biology, Princeton University, Princeton, New Jersey, United States of America. tunstrom@princeton.edu|Department of Ecology and Evolutionary Biology, Princeton University, Princeton, New Jersey, United States of America. tunstrom@princeton.edu|Department of Ecology and Evolutionary Biology, Princeton University, Princeton, New Jersey, United States of America. tunstrom@princeton.edu