Wellcome Trust Centre for Cell Biology, School of Biological Sciences, University of Edinburgh, Edinburgh, EH9?3BF, UK.|European Bioinformatics Institute (EMBL-EBI), European Molecular Biology Laboratory, Wellcome Genome Campus, CB10?1SD, Cambridge, UK.|European Bioinformatics Institute (EMBL-EBI), European Molecular Biology Laboratory, Wellcome Genome Campus, CB10?1SD, Cambridge, UK.|Cambridge Systems Biology Centre, University of Cambridge, Cambridge, CB2xe2x80x891QR, UK.|European Bioinformatics Institute (EMBL-EBI), European Molecular Biology Laboratory, Wellcome Genome Campus, CB10?1SD, Cambridge, UK.|Cambridge Systems Biology Centre, University of Cambridge, Cambridge, CB2xe2x80x891QR, UK.|European Bioinformatics Institute (EMBL-EBI), European Molecular Biology Laboratory, Wellcome Genome Campus, CB10?1SD, Cambridge, UK.|Wellcome Trust Centre for Cell Biology, School of Biological Sciences, University of Edinburgh, Edinburgh, EH9xe2x80x893BF, UK.