School of Biological Sciences, The University of Edinburgh, Swann Building, Kings Buildings, Edinburgh, UK. marco@bioss.ac.uk|School of Biological Sciences, The University of Edinburgh, Swann Building, Kings Buildings, Edinburgh, UK. marco@bioss.ac.uk|School of Biological Sciences, The University of Edinburgh, Swann Building, Kings Buildings, Edinburgh, UK. marco@bioss.ac.uk|School of Biological Sciences, The University of Edinburgh, Swann Building, Kings Buildings, Edinburgh, UK. marco@bioss.ac.uk|School of Biological Sciences, The University of Edinburgh, Swann Building, Kings Buildings, Edinburgh, UK. marco@bioss.ac.uk