Departments of Experimental Plant Ecology and Animal Ecology & Ecophysiology, Radboud University Nijmegen, Nijmegen, The Netherlands, Program for Applied Ecology, Centre for Tropical Forest Science, Smithsonian Tropical Research Institute, Balboa, Ancón, Panamá, Republic of Panamá|Smithsonian Environmental Research Center, Edgewater, Maryland, United States of America.|Smithsonian Environmental Research Center, Edgewater, Maryland, United States of America, Department of Ecology and Evolutionary Biology, University of Connecticut, Storrs, Connecticut, United States of America.|Department of Statistics, Iowa State University, Ames, Iowa, United States of America.|Harvard University, Harvard Forest, Petersham, Massachusetts, United States of America, Bryn Mawr College, Bryn Mawr, Pennsylvania, United States of America.|Departments of Experimental Plant Ecology and Animal Ecology & Ecophysiology, Radboud University Nijmegen, Nijmegen, The Netherlands.