Department of Genetics, Development and Cell Biology, Iowa State University, Ames, IA, 50011, USA.|National Center for Biotechnology Information, National Library of Medicine, National Institutes of Health, Bethesda, MD, 20894, USA.|National Center for Biotechnology Information, National Library of Medicine, National Institutes of Health, Bethesda, MD, 20894, USA.|Department of Biology, Boston University, 5 Cummington Mall, Boston, MA, 02215, USA.|National Center for Biotechnology Information, National Library of Medicine, National Institutes of Health, Bethesda, MD, 20894, USA.|National Center for Biotechnology Information, National Library of Medicine, National Institutes of Health, Bethesda, MD, 20894, USA. spouge@nih.gov.