Department of Ecology and Evolutionary Biology, Princeton University, Princeton, New Jersey, United States of America.|Department of Ecology and Evolutionary Biology, Princeton University, Princeton, New Jersey, United States of America.|Centre for Mathematics and the Environment, University of Exeter, Cornwall, United Kingdom.|Department of Ecology and Evolutionary Biology, Princeton University, Princeton, New Jersey, United States of America.|Department of Ecology and Evolutionary Biology, Princeton University, Princeton, New Jersey, United States of America.