Center for Stock Assessment Research, Department of Applied Mathematics and Statistics, University of California, Santa Cruz, Santa Cruz, California, United States of America, Dipartimento di Elettronica, Informazione e Bioingegneria Politecnico di Milano, Milan, Italy.|Center for Stock Assessment Research, Department of Applied Mathematics and Statistics, University of California, Santa Cruz, Santa Cruz, California, United States of America, Department of Biology, University of Bergen, Bergen, Norway.|Station Biologique de la Tour du Valat, Le Sambuc, Arles, France.|Fisheries Ecology Division, Southwest Fisheries Science Center, National Marine Fisheries Service, NOAA, Santa Cruz, Santa Cruz, California, United States of America.|Department of Mathematics, University of Bergen, Bergen, Norway.