Department of Molecular, Cellular, and Developmental Biology, Yale University, New Haven, Connecticut, United States of America.|Department of Molecular, Cellular, and Developmental Biology, Yale University, New Haven, Connecticut, United States of America.|Department of Molecular, Cellular, and Developmental Biology, Yale University, New Haven, Connecticut, United States of America.|Department of Biochemistry, University of Wisconsin-Madison, Madison, Wisconsin, United States of America.|Department of Molecular, Cellular, and Developmental Biology, Yale University, New Haven, Connecticut, United States of America.