Department of Physics, Chemistry and Biology, Division of Theoretical Biology, Linköping University, Linköping, Sweden.|Department of Biology, Colorado State University, Fort Collins, CO, United States of America.|Zeeman Institute (SBIDER), School of Life Sciences and Mathematics Institute, University of Warwick, Gibbet Hill Road, Coventry, CV4 7AL, UK.|Department of Physics, Chemistry and Biology, Division of Theoretical Biology, Linköping University, Linköping, Sweden.|Department of Biology, Colorado State University, Fort Collins, CO, United States of America.|Department of Physics, Chemistry and Biology, Division of Theoretical Biology, Linköping University, Linköping, Sweden.|Zeeman Institute (SBIDER), School of Life Sciences and Mathematics Institute, University of Warwick, Gibbet Hill Road, Coventry, CV4 7AL, UK.|Department of Physics, Chemistry and Biology, Division of Theoretical Biology, Linköping University, Linköping, Sweden.