Department of Physics and Astronomy, Purdue University, West Lafayette, Indiana, United States of America.|Department of Biophysics, University of Texas Southwestern Medical Center, Dallas, Texas, United States of America.|Department of Physics and Astronomy, Purdue University, West Lafayette, Indiana, United States of America.|Division of Biological Sciences, University of California San Diego, California, United States of America.|Department of Applied Physics and Applied Mathematics, Columbia University, New York, New York, United States of America.|Division of Biological Sciences, University of California San Diego, California, United States of America.|Laboratoire de Physique Théorique, CNRS, Université Pierre et Marie Curie and École Normale Supérieure, Paris, France.