Department of Cell Biology, UCL Institute of Ophthalmology, Bath St, London EC1V 9EL, UK. tim.levine@ucl.ac.uk|Department of Cell Biology, UCL Institute of Ophthalmology, Bath St, London EC1V 9EL, UK. tim.levine@ucl.ac.uk|Department of Cell Biology, UCL Institute of Ophthalmology, Bath St, London EC1V 9EL, UK. tim.levine@ucl.ac.uk|Department of Cell Biology, UCL Institute of Ophthalmology, Bath St, London EC1V 9EL, UK. tim.levine@ucl.ac.uk|Department of Cell Biology, UCL Institute of Ophthalmology, Bath St, London EC1V 9EL, UK. tim.levine@ucl.ac.uk