School of Agriculture and Biology, Shanghai Jiaotong University, Shanghai 200240, China. runqingyang@sjtu.edu.cn|School of Agriculture and Biology, Shanghai Jiaotong University, Shanghai 200240, China. runqingyang@sjtu.edu.cn|School of Agriculture and Biology, Shanghai Jiaotong University, Shanghai 200240, China. runqingyang@sjtu.edu.cn|School of Agriculture and Biology, Shanghai Jiaotong University, Shanghai 200240, China. runqingyang@sjtu.edu.cn|School of Agriculture and Biology, Shanghai Jiaotong University, Shanghai 200240, China. runqingyang@sjtu.edu.cn