Department of Entomology and the Center For Infectious Disease Dynamics, The Pennsylvania State University, University Park, Pennsylvania, United States of America.|Department of Ecology & Evolutionary Biology, University of Toronto, Toronto, Ontario, Canada.|Department of Entomology and the Center For Infectious Disease Dynamics, The Pennsylvania State University, University Park, Pennsylvania, United States of America.|Department of Entomology and the Center For Infectious Disease Dynamics, The Pennsylvania State University, University Park, Pennsylvania, United States of America.