Department of Biology, Emory University, Atlanta, GA 30322, USA.|Department of Biology, Emory University, Atlanta, GA 30322, USA.|Department of Biology, Emory University, Atlanta, GA 30322, USA.|Department of Biology, Emory University, Atlanta, GA 30322, USA.|Department of Biology, Emory University, Atlanta, GA 30322, USA.|Department of Biology, Emory University, Atlanta, GA 30322, USA.