Division of Evolution & Genomic Sciences, University of Manchester, Manchester, MUK.|Wellcome Centre for Cell-Matrix Research, Division of Cell-Matrix Biology and Regenerative Medicine, Faculty of Biology Medicine and Health, University of Manchester, Manchester, MUK.|Wellcome Centre for Cell-Matrix Research, Division of Cell-Matrix Biology and Regenerative Medicine, Faculty of Biology Medicine and Health, University of Manchester, Manchester, MUK.|Division of Evolution & Genomic Sciences, University of Manchester, Manchester, MUK.