Institute of Arable Crop Research, Long Ashton, Bristol, BS41 9AF, UK.|Institute of Arable Crop Research, Long Ashton, Bristol, BS41 9AF, UK.|Institute of Arable Crop Research, Long Ashton, Bristol, BS41 9AF, UK.|Institute of Arable Crop Research, Long Ashton, Bristol, BS41 9AF, UK.|Institute of Arable Crop Research, Long Ashton, Bristol, BS41 9AF, UK.