The Sainsbury Laboratory, John Innes Centre, Colney Lane, Norwich NR47UH, UK. dan.maclean@sainsbury-laboratory.ac.uk|The Sainsbury Laboratory, John Innes Centre, Colney Lane, Norwich NR47UH, UK. dan.maclean@sainsbury-laboratory.ac.uk|The Sainsbury Laboratory, John Innes Centre, Colney Lane, Norwich NR47UH, UK. dan.maclean@sainsbury-laboratory.ac.uk