Division of Computational Biology, School of Life Sciences, University of Dundee, Dundee DD1 4HN, UK.|Division of Computational Biology, School of Life Sciences, University of Dundee, Dundee DD1 4HN, UK.|Division of Computational Biology, School of Life Sciences, University of Dundee, Dundee DD1 4HN, UK.|Division of Computational Biology, School of Life Sciences, University of Dundee, Dundee DD1 4HN, UK.|Division of Computational Biology, School of Life Sciences, University of Dundee, Dundee DD1 4HN, UK.|Division of Computational Biology, School of Life Sciences, University of Dundee, Dundee DD1 4HN, UK.