Department of Biology, Penn State University, University Park, PA 16802, USA. wuc114@psu.edu|Department of Biology, Penn State University, University Park, PA 16802, USA. wuc114@psu.edu|Department of Biology, Penn State University, University Park, PA 16802, USA. wuc114@psu.edu|Department of Biology, Penn State University, University Park, PA 16802, USA. wuc114@psu.edu|Department of Biology, Penn State University, University Park, PA 16802, USA. wuc114@psu.edu