Department of Zoology, University of Oxford, Oxford, OX1 3PS, UK. simon.dellicour@rega.kuleuven.be.|Department of Zoology, University of Oxford, Oxford, OX1 3PS, UK. rebeccarosetm@gmail.com.|Department of Zoology, University of Oxford, Oxford, OX1 3PS, UK. oliver.pybus@zoo.ox.ac.uk.