Computational and Molecular Population Genetics, Zoological Institute, University of Bern, Baltzerstrasse 6, Bern CH-3012, Switzerland. guillaume.laval@zoo.unibe.ch|Computational and Molecular Population Genetics, Zoological Institute, University of Bern, Baltzerstrasse 6, Bern CH-3012, Switzerland. guillaume.laval@zoo.unibe.ch