Department of Ecology and Evolutionary Biology, University of Tennessee, Knoxville, TN 37996-1610, USA. kryberg@utk.edu|Department of Ecology and Evolutionary Biology, University of Tennessee, Knoxville, TN 37996-1610, USA. kryberg@utk.edu|Department of Ecology and Evolutionary Biology, University of Tennessee, Knoxville, TN 37996-1610, USA. kryberg@utk.edu