Department of Biological Sciences, University of Essex, Wivenhoe Park, Colchester, Essex, UK. harry@essex.ac.uk|Department of Biological Sciences, University of Essex, Wivenhoe Park, Colchester, Essex, UK. harry@essex.ac.uk|Department of Biological Sciences, University of Essex, Wivenhoe Park, Colchester, Essex, UK. harry@essex.ac.uk