School of Agriculture and Biology, Shanghai Jiaotong University, Shanghai, PR China. runqingyang@sjtu.edu.cn|School of Agriculture and Biology, Shanghai Jiaotong University, Shanghai, PR China. runqingyang@sjtu.edu.cn|School of Agriculture and Biology, Shanghai Jiaotong University, Shanghai, PR China. runqingyang@sjtu.edu.cn|School of Agriculture and Biology, Shanghai Jiaotong University, Shanghai, PR China. runqingyang@sjtu.edu.cn