Department of Molecular, Cell and Developmental Biology, Yale University, New Haven, CT 06520, USA.|Department of Molecular, Cell and Developmental Biology, Yale University, New Haven, CT 06520, USA.|Department of Molecular, Cell and Developmental Biology, Yale University, New Haven, CT 06520, USA.|Department of Molecular, Cell and Developmental Biology, Yale University, New Haven, CT 06520, USA.|Department of Molecular, Cell and Developmental Biology, Yale University, New Haven, CT 06520, USA.