Department of Mathematics, University College London, Gower Street, London WC1E 6BT, UK.|Department of Mathematics, University College London, Gower Street, London WC1E 6BT, UK.|The Francis Crick Institute, 1 Midland Road, London, NW1 1AT, UK.|Department of Mathematics, University College London, Gower Street, London WC1E 6BT, UK.