Department of Cell Biology and Neuroscience, University of California, Riverside, Riverside, California, United States of America.|Department of Cell Biology and Neuroscience, University of California, Riverside, Riverside, California, United States of America.|Department of Biological Sciences and Bioengineering, Indian Institute of Technology Kanpur, Kanpur, India.|US National Institutes of Health, National Institute of Child Health and Human Development, Bethesda, Maryland, United States of America.|Department of Cell Biology and Neuroscience, University of California, Riverside, Riverside, California, United States of America.