Department of Biology, Duke University, Durham, North Carolina, United States of America. oliver.ratmann@imperial.ac.uk|Department of Biology, Duke University, Durham, North Carolina, United States of America. oliver.ratmann@imperial.ac.uk|Department of Biology, Duke University, Durham, North Carolina, United States of America. oliver.ratmann@imperial.ac.uk|Department of Biology, Duke University, Durham, North Carolina, United States of America. oliver.ratmann@imperial.ac.uk|Department of Biology, Duke University, Durham, North Carolina, United States of America. oliver.ratmann@imperial.ac.uk