Research School of Biology, Australian National University, Canberra, ACT 0200 and CSIRO Plant Industry, Black Mountain Laboratories, Canberra, ACT 2601, Australia.|Research School of Biology, Australian National University, Canberra, ACT 0200 and CSIRO Plant Industry, Black Mountain Laboratories, Canberra, ACT 2601, Australia.|Research School of Biology, Australian National University, Canberra, ACT 0200 and CSIRO Plant Industry, Black Mountain Laboratories, Canberra, ACT 2601, Australia.|Research School of Biology, Australian National University, Canberra, ACT 0200 and CSIRO Plant Industry, Black Mountain Laboratories, Canberra, ACT 2601, Australia.