Laboratory of Socio-Ecology and Social Evolution, Zoological Institute, KU Leuven, Leuven, Belgium.|Mechanical Engineering Department, Middle East Technical University, Ankara, Turkey.|Laboratory of Socio-Ecology and Social Evolution, Zoological Institute, KU Leuven, Leuven, Belgium.|IRIDIA-CoDE, Université Libre de Bruxelles, Brussels, Belgium.|Laboratory of Socio-Ecology and Social Evolution, Zoological Institute, KU Leuven, Leuven, Belgium.