Department of Evolutionary Genetics, Max Planck Institute for Evolutionary Biology, August-Thienemann-Str. 2, Plön, 24306, Germany. dutheil@evolbio.mpg.de.|Institut des Sciences de lÉvolution - Montpellier, Place Eugène Bataillon - C.C. 065 -, Montpellier, 34095, France. emeric.figuet@univ-montp2.fr.