Department of Biology, University of North Carolina, Chapel Hill, NC 27599-3280, USA, Department of Biology, University of North Carolina, Chapel Hill, NC 27599-32800, USA, Department of Microbiology and Immunology, University of North Carolina, Chapel Hill, NC 27599-7290, USA and Curriculum of Genetics and Molecular Biology, University of North Carolina, Chapel Hill, NC, USA Department of Biology, University of North Carolina, Chapel Hill, NC 27599-3280, USA, Department of Biology, University of North Carolina, Chapel Hill, NC 27599-32800, USA, Department of Microbiology and Immunology, University of North Carolina, Chapel Hill, NC 27599-7290, USA and Curriculum of Genetics and Molecular Biology, University of North Carolina, Chapel Hill, NC, USA Department of Biology, University of North Carolina, Chapel Hill, NC 27599-3280, USA, Department of Biology, University of North Carolina, Chapel Hill, NC 27599-32800, USA, Department of Microbiology and Immunology, University of North Carolina, Chapel Hill, NC 27599-7290, USA and Curriculum of Genetics and Molecular Biology, University of North Carolina, Chapel Hill, NC, USA.