Department of Biology, San Diego State University, San Diego, CA, USA. yyu@rohan.sdsu.edu|Department of Biology, San Diego State University, San Diego, CA, USA. yyu@rohan.sdsu.edu|Department of Biology, San Diego State University, San Diego, CA, USA. yyu@rohan.sdsu.edu|Department of Biology, San Diego State University, San Diego, CA, USA. yyu@rohan.sdsu.edu