Key Laboratory of Ornamental Plants Germplasm Innovation and Molecular Breeding, National Engineering Research Center for Floriculture, Beijing Forestry University, Beijing, China.|Center for Computational Biology, College of Biological Sciences and Technology, Beijing Forestry University, Beijing, China.|Center for Computational Biology, College of Biological Sciences and Technology, Beijing Forestry University, Beijing, China.|Center for Computational Biology, College of Biological Sciences and Technology, Beijing Forestry University, Beijing, China.|Statistical Genetics, Department of Public Health Sciences, Pennsylvania State University, Pennsylvania, USA.|Center for Computational Biology, College of Biological Sciences and Technology, Beijing Forestry University, Beijing, China.|Center for Computational Biology, College of Biological Sciences and Technology, Beijing Forestry University, Beijing, China.|Ornamental Genetics, National Engineering Research, Center for Floriculture, Beijing Forestry University Beijing, China.|Ornamental Genetics, National Engineering Research, Center for Floriculture, Beijing Forestry University Beijing, China.|Center for Computational Biology, College of Biological Sciences and Technology, Beijing Forestry University, Beijing, China.