Laboratory for Neural Circuit Theory, RIKEN Brain Science Institute, Wako, Saitama, Japan. yasuhirotsubo@riken.jp|Laboratory for Neural Circuit Theory, RIKEN Brain Science Institute, Wako, Saitama, Japan. yasuhirotsubo@riken.jp|Laboratory for Neural Circuit Theory, RIKEN Brain Science Institute, Wako, Saitama, Japan. yasuhirotsubo@riken.jp