Department of Biology, University of Waterloo, Waterloo, Ontario N2L 3G1, Canada.|Department of Biology, University of Waterloo, Waterloo, Ontario N2L 3G1, Canada.|Department of Biology, University of Waterloo, Waterloo, Ontario N2L 3G1, Canada.|Department of Biology, University of Waterloo, Waterloo, Ontario N2L 3G1, Canada.