Centre for Crop Systems Analysis, Wageningen University, Wageningen, The Netherlands.|Centre for Crop Systems Analysis, Wageningen University, Wageningen, The Netherlands.|Theoretical Biology, Kyushu University, Fukuoka, Japan.|Plant Ecophysiology, Utrecht University, Utrecht, the Netherlands.|Centre for Crop Systems Analysis, Wageningen University, Wageningen, The Netherlands.|Centre for Crop Systems Analysis, Wageningen University, Wageningen, The Netherlands.