Department of Public Health Science, Seoul National University, Seoul, Korea. won1@snu.ac.kr.|Interdisciplinary Program of Bioinformatics, Seoul National University, Seoul, Korea. dnjswlzz11@gmail.com.|Interdisciplinary Program of Bioinformatics, Seoul National University, Seoul, Korea. biznok@gmail.com.|The Center for Genome Science, Korea National Institute of Health, KCDC, Osong, Korea. lyou7688@gmail.com.|Department of Public Health Science, Seoul National University, Seoul, Korea. jsung@snu.ac.kr.|Interdisciplinary Program of Bioinformatics, Seoul National University, Seoul, Korea. tspark@stats.snu.ac.kr.