Wageningen UR Plant Breeding, Wageningen University & Research Centre, PO Box 6700 AJ, Wageningen, The Netherlands.|Wageningen UR Plant Breeding, Wageningen University & Research Centre, PO Box 6700 AJ, Wageningen, The Netherlands.|Wageningen UR Plant Breeding, Wageningen University & Research Centre, PO Box 6700 AJ, Wageningen, The Netherlands.|Wageningen UR Plant Breeding, Wageningen University & Research Centre, PO Box 6700 AJ, Wageningen, The Netherlands. chris.maliepaard@wur.nl.