Department of Zoology, University of Oxford, Oxford, United Kingdom. philippe.lemey@zoo.ox.ac.uk|Department of Zoology, University of Oxford, Oxford, United Kingdom. philippe.lemey@zoo.ox.ac.uk|Department of Zoology, University of Oxford, Oxford, United Kingdom. philippe.lemey@zoo.ox.ac.uk|Department of Zoology, University of Oxford, Oxford, United Kingdom. philippe.lemey@zoo.ox.ac.uk|Department of Zoology, University of Oxford, Oxford, United Kingdom. philippe.lemey@zoo.ox.ac.uk|Department of Zoology, University of Oxford, Oxford, United Kingdom. philippe.lemey@zoo.ox.ac.uk|Department of Zoology, University of Oxford, Oxford, United Kingdom. philippe.lemey@zoo.ox.ac.uk|Department of Zoology, University of Oxford, Oxford, United Kingdom. philippe.lemey@zoo.ox.ac.uk