Institute of Biological, Environmental, and Rural Sciences (IBERS), Aberystwyth University, Penglais, Aberystwyth, Ceredigion, SY23 3FG, UK.|Institute of Biological, Environmental, and Rural Sciences (IBERS), Aberystwyth University, Penglais, Aberystwyth, Ceredigion, SY23 3FG, UK.|Institute of Biological, Environmental, and Rural Sciences (IBERS), Aberystwyth University, Penglais, Aberystwyth, Ceredigion, SY23 3FG, UK.|Institute of Biological, Environmental, and Rural Sciences (IBERS), Aberystwyth University, Penglais, Aberystwyth, Ceredigion, SY23 3FG, UK.|Institute of Biological, Environmental, and Rural Sciences (IBERS), Aberystwyth University, Penglais, Aberystwyth, Ceredigion, SY23 3FG, UK. mts11@aber.ac.uk.