Department of Developmental Biology, The Roslin Institute, University of Edinburgh, Easter Bush Campus, Midlothian EH25 9RG, UK.|Department of Developmental Biology, The Roslin Institute, University of Edinburgh, Easter Bush Campus, Midlothian EH25 9RG, UK.|Department of Developmental Biology, The Roslin Institute, University of Edinburgh, Easter Bush Campus, Midlothian EH25 9RG, UK.