School of Mathematics and Institute of Molecular and Cellular Biology, University of Leeds, Leeds LS2 9JT, UK.|School of Mathematics and Institute of Molecular and Cellular Biology, University of Leeds, Leeds LS2 9JT, UK.|School of Mathematics and Institute of Molecular and Cellular Biology, University of Leeds, Leeds LS2 9JT, UK.|School of Mathematics and Institute of Molecular and Cellular Biology, University of Leeds, Leeds LS2 9JT, UK.