University of Colorado Museum of Natural History and Department of Ecology and Evolutionary Biology, University of Colorado Boulder, Boulder, CO 80309-0265, USA. Andrew.Hill@colorado.edu|University of Colorado Museum of Natural History and Department of Ecology and Evolutionary Biology, University of Colorado Boulder, Boulder, CO 80309-0265, USA. Andrew.Hill@colorado.edu|University of Colorado Museum of Natural History and Department of Ecology and Evolutionary Biology, University of Colorado Boulder, Boulder, CO 80309-0265, USA. Andrew.Hill@colorado.edu|University of Colorado Museum of Natural History and Department of Ecology and Evolutionary Biology, University of Colorado Boulder, Boulder, CO 80309-0265, USA. Andrew.Hill@colorado.edu|University of Colorado Museum of Natural History and Department of Ecology and Evolutionary Biology, University of Colorado Boulder, Boulder, CO 80309-0265, USA. Andrew.Hill@colorado.edu|University of Colorado Museum of Natural History and Department of Ecology and Evolutionary Biology, University of Colorado Boulder, Boulder, CO 80309-0265, USA. Andrew.Hill@colorado.edu|University of Colorado Museum of Natural History and Department of Ecology and Evolutionary Biology, University of Colorado Boulder, Boulder, CO 80309-0265, USA. Andrew.Hill@colorado.edu|University of Colorado Museum of Natural History and Department of Ecology and Evolutionary Biology, University of Colorado Boulder, Boulder, CO 80309-0265, USA. Andrew.Hill@colorado.edu