Division of Computational Biology, School of Life Sciences, University of Dundee, Dundee, DD1 5EH, UK.|Division of Computational Biology, School of Life Sciences, University of Dundee, Dundee, DD1 5EH, UK.|Molecular Medicine, School of Life Sciences, University of Dundee, Dundee, DD1 5RH, UK.|Division of Computational Biology, School of Life Sciences, University of Dundee, Dundee, DD1 5EH, UK.