Department of Developmental and Cell Biology, University of California, Irvine, California, United States of America , Center for Complex Biological Systems, University of California, Irvine, California, United States of America , Department of Computer Science, University of California, Irvine, California, United States of America.|Department of Developmental and Cell Biology, University of California, Irvine, California, United States of America.|Department of Developmental and Cell Biology, University of California, Irvine, California, United States of America , Department of Developmental Neurobiology, St Jude Childrens Research Hospital, Memphis, Tennessee, United States of America.|Department of Pathology and Laboratory Medicine, University of California, Irvine, California, United States of America.|Department of Computer Science, University of California, Irvine, California, United States of America.|Department of Developmental and Cell Biology, University of California, Irvine, California, United States of America , Center for Complex Biological Systems, University of California, Irvine, California, United States of America.|Department of Developmental and Cell Biology, University of California, Irvine, California, United States of America , Center for Complex Biological Systems, University of California, Irvine, California, United States of America , Department of Pathology and Laboratory Medicine, University of California, Irvine, California, United States of America.