Department of Biological Sciences, University of Essex, Wivenhoe Park, Colchester, Essex CO4 3SQ, UK. mastal@essex.ac.uk <mastal@essex.ac.uk>|Department of Biological Sciences, University of Essex, Wivenhoe Park, Colchester, Essex CO4 3SQ, UK. mastal@essex.ac.uk <mastal@essex.ac.uk>