Department of Biological Sciences, University of Calgary, AB, T2N 1N4 Canada. debridge@ucalgary.ca <debridge@ucalgary.ca>|Department of Biological Sciences, University of Calgary, AB, T2N 1N4 Canada. debridge@ucalgary.ca <debridge@ucalgary.ca>|Department of Biological Sciences, University of Calgary, AB, T2N 1N4 Canada. debridge@ucalgary.ca <debridge@ucalgary.ca>