Department of Ecology and Evolutionary Biology, University of Tennessee, Knoxville, TN 37996, USA. mikeg@utk.edu|Department of Ecology and Evolutionary Biology, University of Tennessee, Knoxville, TN 37996, USA. mikeg@utk.edu|Department of Ecology and Evolutionary Biology, University of Tennessee, Knoxville, TN 37996, USA. mikeg@utk.edu