Division of Molecular Biosciences, Faculty of Natural Sciences, Imperial College London, London, United Kingdom|Division of Molecular Biosciences, Faculty of Natural Sciences, Imperial College London, London, United Kingdom|Division of Molecular Biosciences, Faculty of Natural Sciences, Imperial College London, London, United Kingdom|Division of Molecular Biosciences, Faculty of Natural Sciences, Imperial College London, London, United Kingdom|Division of Molecular Biosciences, Faculty of Natural Sciences, Imperial College London, London, United Kingdom