School of Animal and Microbial Sciences, University of Reading Whiteknights, PO Box 228, Reading RG6 6AJ, UK.|School of Animal and Microbial Sciences, University of Reading Whiteknights, PO Box 228, Reading RG6 6AJ, UK.|School of Animal and Microbial Sciences, University of Reading Whiteknights, PO Box 228, Reading RG6 6AJ, UK.