Institute of Evolutionary and Ecological Sciences, Kaiserstraat 63, Leiden University 2311 GP Leiden, The Netherlands. wpiel@buffalo.edu|Institute of Evolutionary and Ecological Sciences, Kaiserstraat 63, Leiden University 2311 GP Leiden, The Netherlands. wpiel@buffalo.edu|Institute of Evolutionary and Ecological Sciences, Kaiserstraat 63, Leiden University 2311 GP Leiden, The Netherlands. wpiel@buffalo.edu