Biotechnology Research Institute, Chinese Academy of Agricultural Sciences, Beijing 100081, PR China. tianjian3721@163.com|Biotechnology Research Institute, Chinese Academy of Agricultural Sciences, Beijing 100081, PR China. tianjian3721@163.com|Biotechnology Research Institute, Chinese Academy of Agricultural Sciences, Beijing 100081, PR China. tianjian3721@163.com|Biotechnology Research Institute, Chinese Academy of Agricultural Sciences, Beijing 100081, PR China. tianjian3721@163.com