Center for Quantitative Biology, Academy for Advanced Interdisciplinary Studies, Peking University, Beijing, China.|The State Key Laboratory for Artificial Microstructures and Mesoscopic Physics, School of Physics, Peking University, Beijing, China.|Center for Quantitative Biology, Academy for Advanced Interdisciplinary Studies, Peking University, Beijing, China.|School of Informatics and Computing, Indiana University, Bloomington, Indiana, United States of America.|Center for Quantitative Biology, Academy for Advanced Interdisciplinary Studies, Peking University, Beijing, China.|Center for Quantitative Biology, Academy for Advanced Interdisciplinary Studies, Peking University, Beijing, China.|Section of Molecular Biology, Division of Biological Sciences, University of California San Diego, La Jolla, California, United States of America.|Center for Quantitative Biology, Academy for Advanced Interdisciplinary Studies, Peking University, Beijing, China.