Program for Evolutionary Dynamics, Harvard University, Cambridge, Massachusetts, United States of America. niko.beerenwinkel@bsse.ethz.ch|Program for Evolutionary Dynamics, Harvard University, Cambridge, Massachusetts, United States of America. niko.beerenwinkel@bsse.ethz.ch|Program for Evolutionary Dynamics, Harvard University, Cambridge, Massachusetts, United States of America. niko.beerenwinkel@bsse.ethz.ch|Program for Evolutionary Dynamics, Harvard University, Cambridge, Massachusetts, United States of America. niko.beerenwinkel@bsse.ethz.ch|Program for Evolutionary Dynamics, Harvard University, Cambridge, Massachusetts, United States of America. niko.beerenwinkel@bsse.ethz.ch|Program for Evolutionary Dynamics, Harvard University, Cambridge, Massachusetts, United States of America. niko.beerenwinkel@bsse.ethz.ch|Program for Evolutionary Dynamics, Harvard University, Cambridge, Massachusetts, United States of America. niko.beerenwinkel@bsse.ethz.ch|Program for Evolutionary Dynamics, Harvard University, Cambridge, Massachusetts, United States of America. niko.beerenwinkel@bsse.ethz.ch