Department of Microbiology and Immunology, Columbia University College of Physicians and Surgeons, New York, New York, United States of America , School of International and Public Affairs, Columbia University, New York, New York, United States of America , Bloomberg School of Public Health, John Hopkins University, Baltimore, Maryland, United States of America.|Spatial Ecology and Epidemiology Group, Department of Zoology, University of Oxford, Oxford, United Kingdom.|Spatial Ecology and Epidemiology Group, Department of Zoology, University of Oxford, Oxford, United Kingdom.|Bloomberg School of Public Health, John Hopkins University, Baltimore, Maryland, United States of America.|Department of Microbiology and Immunology, Columbia University College of Physicians and Surgeons, New York, New York, United States of America , Division of Infectious Diseases, Department of Medicine, Columbia University College of Physicians and Surgeons, New York, New York, United States of America.