Center for Theoretical Biological Physics, University of California San Diego, La Jolla, California, United States of America. hlevine@ucsd.edu|Center for Theoretical Biological Physics, University of California San Diego, La Jolla, California, United States of America. hlevine@ucsd.edu|Center for Theoretical Biological Physics, University of California San Diego, La Jolla, California, United States of America. hlevine@ucsd.edu