Department of Biology, Case Western Reserve University, Cleveland, Ohio, United States of America.|Department of Biology, Case Western Reserve University, Cleveland, Ohio, United States of America.|Department of Biology, Case Western Reserve University, Cleveland, Ohio, United States of America.|Department of Biology, Case Western Reserve University, Cleveland, Ohio, United States of America.|Department of Biology, Case Western Reserve University, Cleveland, Ohio, United States of America, Department of Genetics and Genome Sciences, Case Western Reserve University, Cleveland, Ohio, United States of America.