Research School of Biological Sciences, The Australian National University, GPO Box 475, Canberra ACT 2601, Australia. mgibbs@rsbs.anu.edu.au|Research School of Biological Sciences, The Australian National University, GPO Box 475, Canberra ACT 2601, Australia. mgibbs@rsbs.anu.edu.au|Research School of Biological Sciences, The Australian National University, GPO Box 475, Canberra ACT 2601, Australia. mgibbs@rsbs.anu.edu.au