Computation and Neural Systems Program, California Institute of Technology, Pasadena, CA, United States of America.|Computation and Neural Systems Program, California Institute of Technology, Pasadena, CA, United States of America.|Division of the Humanities and Social Sciences, California Institute of Technology, Pasadena, CA, United States of America.|Division of the Humanities and Social Sciences, California Institute of Technology, Pasadena, CA, United States of America.|Computation and Neural Systems Program, California Institute of Technology, Pasadena, CA, United States of America.