Center for Ecology and Evolutionary Biology, University of Oregon, Eugene, OR, USA.|Center for Ecology and Evolutionary Biology, University of Oregon, Eugene, OR, USA.|Center for Ecology and Evolutionary Biology, University of Oregon, Eugene, OR, USA.|Center for Ecology and Evolutionary Biology, University of Oregon, Eugene, OR, USA.|Center for Ecology and Evolutionary Biology, University of Oregon, Eugene, OR, USA.|Center for Ecology and Evolutionary Biology, University of Oregon, Eugene, OR, USA.|Center for Ecology and Evolutionary Biology, University of Oregon, Eugene, OR, USA.|Center for Ecology and Evolutionary Biology, University of Oregon, Eugene, OR, USA.