Department of Cell and Developmental Biology, University of Michigan Medical School, Ann Arbor, MI, USA.|Department of Cell and Developmental Biology, University of Michigan Medical School, Ann Arbor, MI, USA.|Department of Cell and Developmental Biology, University of Michigan Medical School, Ann Arbor, MI, USA.|Department of Biology, Washington University in St. Louis, St. Louis, MO, USA.|Department of Cell and Developmental Biology, University of Michigan Medical School, Ann Arbor, MI, USA.|Department of Cell and Developmental Biology, University of Michigan Medical School, Ann Arbor, MI, USA.|Department of Neurology, Boston Childrens Hospital, Harvard Medical School, Boston, MA, USA.|Department of Biology, Washington University in St. Louis, St. Louis, MO, USA.|Department of Cell and Developmental Biology, University of Michigan Medical School, Ann Arbor, MI, USA.|Department of Cell and Developmental Biology, University of Michigan Medical School, Ann Arbor, MI, USA.|Department of Cell and Developmental Biology, University of Michigan Medical School, Ann Arbor, MI, USA.|Department of Cell and Developmental Biology, University of Michigan Medical School, Ann Arbor, MI, USA.|Department of Cell and Developmental Biology, University of Michigan Medical School, Ann Arbor, MI, USA.|Department of Biology, University of Toronto, Toronto, ON, Canada.|Department of Cell and Developmental Biology, University of Michigan Medical School, Ann Arbor, MI, USA.|Department of Molecular and Cellular Biology Cambridge, MA, USA.|Department of Neurology, Boston Childrens Hospital, Harvard Medical School, Boston, MA, USA.|Department of Biology, Washington University in St. Louis, St. Louis, MO, USA.|Department of Cell and Developmental Biology, University of Michigan Medical School, Ann Arbor, MI, USA.