Marine Biology Research Division, Scripps Institution of Oceanography University of California at San Diego, La Jolla, CA 92093 USA. spodell@ucsd.edu|Marine Biology Research Division, Scripps Institution of Oceanography University of California at San Diego, La Jolla, CA 92093 USA. spodell@ucsd.edu|Marine Biology Research Division, Scripps Institution of Oceanography University of California at San Diego, La Jolla, CA 92093 USA. spodell@ucsd.edu