Joint Carnegie Mellon/University of Pittsburgh Ph.D. Program in Computational Biology, Lane Center for Computational Biology, Carnegie Mellon University, Pittsburgh, PA 15213, USA. sachowdh@andrew.cmu.edu|Joint Carnegie Mellon/University of Pittsburgh Ph.D. Program in Computational Biology, Lane Center for Computational Biology, Carnegie Mellon University, Pittsburgh, PA 15213, USA. sachowdh@andrew.cmu.edu|Joint Carnegie Mellon/University of Pittsburgh Ph.D. Program in Computational Biology, Lane Center for Computational Biology, Carnegie Mellon University, Pittsburgh, PA 15213, USA. sachowdh@andrew.cmu.edu|Joint Carnegie Mellon/University of Pittsburgh Ph.D. Program in Computational Biology, Lane Center for Computational Biology, Carnegie Mellon University, Pittsburgh, PA 15213, USA. sachowdh@andrew.cmu.edu|Joint Carnegie Mellon/University of Pittsburgh Ph.D. Program in Computational Biology, Lane Center for Computational Biology, Carnegie Mellon University, Pittsburgh, PA 15213, USA. sachowdh@andrew.cmu.edu|Joint Carnegie Mellon/University of Pittsburgh Ph.D. Program in Computational Biology, Lane Center for Computational Biology, Carnegie Mellon University, Pittsburgh, PA 15213, USA. sachowdh@andrew.cmu.edu