Department of Biology, Indiana University, Bloomington, IN 47405, USA.|Department of Biology, Indiana University, Bloomington, IN 47405, USA.|Department of Biology, Indiana University, Bloomington, IN 47405, USA.|Department of Biology, Indiana University, Bloomington, IN 47405, USA.|Department of Biology, Indiana University, Bloomington, IN 47405, USA.|Department of Biology, Indiana University, Bloomington, IN 47405, USA.|Department of Biology, Indiana University, Bloomington, IN 47405, USA.|Department of Biology, Indiana University, Bloomington, IN 47405, USA.|Department of Biology, Indiana University, Bloomington, IN 47405, USA.|Department of Biology, Indiana University, Bloomington, IN 47405, USA.|Department of Biology, Indiana University, Bloomington, IN 47405, USA.|Department of Biology, Indiana University, Bloomington, IN 47405, USA.|Department of Biology, Indiana University, Bloomington, IN 47405, USA.