Division of Biology and Biomedical Sciences, Washington University in St. Louis, One Brookings Drive, Brauer 2004, St. Louis, MO 63130, USA.|Division of Biology and Biomedical Sciences, Washington University in St. Louis, One Brookings Drive, Brauer 2004, St. Louis, MO 63130, USA.|Division of Biology and Biomedical Sciences, Washington University in St. Louis, One Brookings Drive, Brauer 2004, St. Louis, MO 63130, USA.|Division of Biology and Biomedical Sciences, Washington University in St. Louis, One Brookings Drive, Brauer 2004, St. Louis, MO 63130, USA.