Department of Mathematics, Faculty of Natural Sciences, Imperial College, London SW7 2AZ, UK.|Department of Mathematics, Faculty of Natural Sciences, Imperial College, London SW7 2AZ, UK.|Department of Mathematics, Faculty of Natural Sciences, Imperial College, London SW7 2AZ, UK.|Department of Mathematics, Faculty of Natural Sciences, Imperial College, London SW7 2AZ, UK.|MRC London Institute of Medical Sciences (LMS), Du Cane Road, London W12 0NN, UK.|MRC London Institute of Medical Sciences (LMS), Du Cane Road, London W12 0NN, UK.|Department of Mathematics, Faculty of Natural Sciences, Imperial College, London SW7 2AZ, UK.