Department of Ecology and Evolutionary Biology, University of Colorado, Boulder, Colorado, USA. stuckyb@colorado.edu.|Department of Ecology and Evolutionary Biology, University of Colorado, Boulder, Colorado, USA. stuckyb@colorado.edu.|Department of Ecology and Evolutionary Biology, University of Colorado, Boulder, Colorado, USA. stuckyb@colorado.edu.|Department of Ecology and Evolutionary Biology, University of Colorado, Boulder, Colorado, USA. stuckyb@colorado.edu.|Department of Ecology and Evolutionary Biology, University of Colorado, Boulder, Colorado, USA. stuckyb@colorado.edu.|Department of Ecology and Evolutionary Biology, University of Colorado, Boulder, Colorado, USA. stuckyb@colorado.edu.