School of Biology, University of Leeds, Leeds, United Kingdom.|School of Biology, University of Leeds, Leeds, United Kingdom, Center for Advanced Studies (CEAB-CSIC), Blanes, Spain.|Groningen Institute for Evolutionary Life Sciences, University of Groningen, Groningen, Netherlands.|School of Biology, University of Leeds, Leeds, United Kingdom, Institute of Integrative Biology, University of Liverpool, Liverpool, United Kingdom (current address).