Department of Biology, Brigham Young University, Provo, UT, USA.|Department of Biology, Brigham Young University, Provo, UT, USA.|Department of Biology, Brigham Young University, Provo, UT, USA.|Department of Biology, Brigham Young University, Provo, UT, USA.|Department of Biology, Brigham Young University, Provo, UT, USA.|Department of Biology, Brigham Young University, Provo, UT, USA.|Department of Biology, Brigham Young University, Provo, UT, USA.|Department of Biology, Brigham Young University, Provo, UT, USA.|Department of Biology, Brigham Young University, Provo, UT, USA.|Department of Biology, Brigham Young University, Provo, UT, USA. perry.ridge@byu.edu.