Marine Biology and Ecology Research Centre, School of Marine Science and Engineering, Plymouth University, Plymouth, Devon PL4 8AA, UK. oliver.tills@plymouth.ac.uk|Marine Biology and Ecology Research Centre, School of Marine Science and Engineering, Plymouth University, Plymouth, Devon PL4 8AA, UK. oliver.tills@plymouth.ac.uk|Marine Biology and Ecology Research Centre, School of Marine Science and Engineering, Plymouth University, Plymouth, Devon PL4 8AA, UK. oliver.tills@plymouth.ac.uk|Marine Biology and Ecology Research Centre, School of Marine Science and Engineering, Plymouth University, Plymouth, Devon PL4 8AA, UK. oliver.tills@plymouth.ac.uk|Marine Biology and Ecology Research Centre, School of Marine Science and Engineering, Plymouth University, Plymouth, Devon PL4 8AA, UK. oliver.tills@plymouth.ac.uk