Department of Computer Science, University of Oxford, Oxford, United Kingdom.|Department of Computer Science, University of Oxford, Oxford, United Kingdom, Department of Mathematics, Queensland University of Technology, Brisbane, Queensland, Australia.|Department of Applied Mathematics and Statistics, University of California Santa Cruz, Santa Cruz, California, United States of America, Department of Biology, University of Bergen, Bergen, Norway.|Mathematical Ecology Research Group, Department of Zoology, University of Oxford, Oxford, United Kingdom.