BIO5 Institute, School of Plant Sciences, University of Arizona, Tucson, AZ, USA.|BIO5 Institute, School of Plant Sciences, University of Arizona, Tucson, AZ, USA.|BIO5 Institute, School of Plant Sciences, University of Arizona, Tucson, AZ, USA.|Department of Biology, University of Pennsylvania, Philadelphia, PA, USA.|BIO5 Institute, School of Plant Sciences, University of Arizona, Tucson, AZ, USA.