Department of Ecology and Evolutionary Biology, Princeton University, Princeton, New Jersey, USA. sf7@princeton.edu|Department of Ecology and Evolutionary Biology, Princeton University, Princeton, New Jersey, USA. sf7@princeton.edu|Department of Ecology and Evolutionary Biology, Princeton University, Princeton, New Jersey, USA. sf7@princeton.edu|Department of Ecology and Evolutionary Biology, Princeton University, Princeton, New Jersey, USA. sf7@princeton.edu|Department of Ecology and Evolutionary Biology, Princeton University, Princeton, New Jersey, USA. sf7@princeton.edu