Institute of Biomedical Sciences, Academia Sinica, Taipei, Taiwan. wliu56@gate.sinica.edu.tw|Institute of Biomedical Sciences, Academia Sinica, Taipei, Taiwan. wliu56@gate.sinica.edu.tw|Institute of Biomedical Sciences, Academia Sinica, Taipei, Taiwan. wliu56@gate.sinica.edu.tw|Institute of Biomedical Sciences, Academia Sinica, Taipei, Taiwan. wliu56@gate.sinica.edu.tw|Institute of Biomedical Sciences, Academia Sinica, Taipei, Taiwan. wliu56@gate.sinica.edu.tw|Institute of Biomedical Sciences, Academia Sinica, Taipei, Taiwan. wliu56@gate.sinica.edu.tw