Program for Neural Computation, Carnegie Mellon University, Pittsburgh, Pennsylvania, United States of America, Center for the Neural Basis of Cognition, Pittsburgh, Pennsylvania, United States of America.|Center for the Neural Basis of Cognition, Pittsburgh, Pennsylvania, United States of America, Department of Biological Sciences, Carnegie Mellon University, Pittsburgh, Pennsylvania, United States of America.|Center for the Neural Basis of Cognition, Pittsburgh, Pennsylvania, United States of America, Department of Ophthalmology, University of Pittsburgh, Pittsburgh, Pennsylvania, United States of America.|Center for the Neural Basis of Cognition, Pittsburgh, Pennsylvania, United States of America, Department of Ophthalmology, University of Pittsburgh, Pittsburgh, Pennsylvania, United States of America.|Center for the Neural Basis of Cognition, Pittsburgh, Pennsylvania, United States of America, Department of Biological Sciences, Carnegie Mellon University, Pittsburgh, Pennsylvania, United States of America, Department of Neurobiology, University of Pittsburgh, Pittsburgh, Pennsylvania, United States of America.|Center for the Neural Basis of Cognition, Pittsburgh, Pennsylvania, United States of America, Department of Statistics, Carnegie Mellon University, Pittsburgh, Pennsylvania, United States of America, Department of Machine Learning, Carnegie Mellon University, Pittsburgh, Pennsylvania, United States of America.