ARC Centre of Excellence in Vision Science, Australian National University, Acton, Canberra, Australian Capital Territory, Australia, Research School of Biology, Australian National University, Acton, Canberra, Australian Capital Territory, Australia, Queensland Brain Institute, The University of Queensland, St Lucia, Brisbane, Queensland, Australia.|Queensland Brain Institute, The University of Queensland, St Lucia, Brisbane, Queensland, Australia.|ARC Centre of Excellence in Vision Science, Australian National University, Acton, Canberra, Australian Capital Territory, Australia, National Vision Research Institute, Australian College of Optometry, Carlton, Melbourne, Victoria, Australia.|ARC Centre of Excellence in Vision Science, Australian National University, Acton, Canberra, Australian Capital Territory, Australia, Queensland Brain Institute, The University of Queensland, St Lucia, Brisbane, Queensland, Australia, School of Information Technology and Electrical Engineering, The University of Queensland, St Lucia, Brisbane, Queensland, Australia.